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  <front>
    <journal-meta>
      <journal-id journal-id-type="publisher-id">52</journal-id>
      <journal-id journal-id-type="index">urn:lsid:arphahub.com:pub:4A25E796-D4BE-573E-A5A5-2A05A815D1C2</journal-id>
      <journal-title-group>
        <journal-title xml:lang="en">One Ecosystem</journal-title>
        <abbrev-journal-title xml:lang="en">OE</abbrev-journal-title>
      </journal-title-group>
      <issn pub-type="epub">2367-8194</issn>
      <publisher>
        <publisher-name>Pensoft Publishers</publisher-name>
      </publisher>
    </journal-meta>
    <article-meta>
      <article-id pub-id-type="doi">10.3897/oneeco.11.e189660</article-id>
      <article-id pub-id-type="publisher-id">189660</article-id>
      <article-id pub-id-type="manuscript">30303</article-id>
      <article-categories>
        <subj-group subj-group-type="heading">
          <subject>Research Article</subject>
        </subj-group>
        <subj-group subj-group-type="scientific_subject">
          <subject>Ecological expertise &amp; Expert systems</subject>
          <subject>Ecology &amp; Environmental sciences</subject>
          <subject>Habitats</subject>
          <subject> Ecosystems &amp; Natural Spaces</subject>
        </subj-group>
        <subj-group subj-group-type="geographical_area">
          <subject>Black Sea</subject>
        </subj-group>
      </article-categories>
      <title-group>
        <article-title>Development and application of the Eco-Geomorphological Dune Value Index (EGDVI) for assessment of conservation status: a pilot case at the Strandzha Black Sea Coast, Bulgaria</article-title>
      </title-group>
      <contrib-group content-type="authors">
        <contrib contrib-type="author" corresp="yes">
          <name name-style="western">
            <surname>Prodanov</surname>
            <given-names>Bogdan</given-names>
          </name>
          <email xlink:type="simple">bogdanprodanov@gmail.com</email>
          <uri content-type="orcid">https://orcid.org/0000-0002-8118-3034</uri>
          <xref ref-type="aff" rid="A1">1</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Bekova</surname>
            <given-names>Radoslava</given-names>
          </name>
          <uri content-type="orcid">https://orcid.org/0000-0002-9353-1956</uri>
          <xref ref-type="aff" rid="A1">1</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Valcheva</surname>
            <given-names>Magdalena</given-names>
          </name>
          <uri content-type="orcid">https://orcid.org/0000-0003-4327-6501</uri>
          <xref ref-type="aff" rid="A2">2</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Tzonev</surname>
            <given-names>Rossen</given-names>
          </name>
          <uri content-type="orcid">https://orcid.org/0000-0001-8112-1354</uri>
          <xref ref-type="aff" rid="A3">3</xref>
        </contrib>
        <contrib contrib-type="author" corresp="no">
          <name name-style="western">
            <surname>Baltakova</surname>
            <given-names>Ahinora</given-names>
          </name>
          <uri content-type="orcid">https://orcid.org/0000-0001-6973-9688</uri>
          <xref ref-type="aff" rid="A3">3</xref>
        </contrib>
      </contrib-group>
      <aff id="A1">
        <label>1</label>
        <addr-line content-type="verbatim">Institute of Oceanology, Bulgarian Academy of Sciences, Varna, Bulgaria</addr-line>
        <institution>Institute of Oceanology, Bulgarian Academy of Sciences</institution>
        <addr-line content-type="city">Varna</addr-line>
        <country>Bulgaria</country>
        <uri content-type="ror">https://ror.org/01x8hew03</uri>
      </aff>
      <aff id="A2">
        <label>2</label>
        <addr-line content-type="verbatim">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, Sofia, Bulgaria</addr-line>
        <institution>Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences</institution>
        <addr-line content-type="city">Sofia</addr-line>
        <country>Bulgaria</country>
        <uri content-type="ror">https://ror.org/01x8hew03</uri>
      </aff>
      <aff id="A3">
        <label>3</label>
        <addr-line content-type="verbatim">Sofia University “St. Kliment Ohridski”, Sofia, Bulgaria</addr-line>
        <institution>Sofia University “St. Kliment Ohridski”</institution>
        <addr-line content-type="city">Sofia</addr-line>
        <country>Bulgaria</country>
        <uri content-type="ror">https://ror.org/02jv3k292</uri>
      </aff>
      <author-notes>
        <fn fn-type="corresp">
          <p>Corresponding author: Bogdan Prodanov (<email xlink:type="simple">bogdanprodanov@gmail.com</email>).</p>
        </fn>
        <fn fn-type="edited-by">
          <p>Academic editor: Joachim Maes</p>
        </fn>
      </author-notes>
      <pub-date pub-type="collection">
        <year>2026</year>
      </pub-date>
      <pub-date pub-type="epub">
        <day>22</day>
        <month>05</month>
        <year>2026</year>
      </pub-date>
      <volume>11</volume>
      <elocation-id>e189660</elocation-id>
      <uri content-type="arpha" xlink:href="http://openbiodiv.net/31FDE319-322D-5D34-B502-D28AFB205F26">31FDE319-322D-5D34-B502-D28AFB205F26</uri>
      <history>
        <date date-type="received">
          <day>24</day>
          <month>02</month>
          <year>2026</year>
        </date>
        <date date-type="accepted">
          <day>11</day>
          <month>05</month>
          <year>2026</year>
        </date>
      </history>
      <permissions>
        <copyright-statement>Bogdan Prodanov, Radoslava Bekova, Magdalena Valcheva, Rossen Tzonev, Ahinora Baltakova</copyright-statement>
        <license license-type="creative-commons-attribution" xlink:href="http://creativecommons.org/licenses/by/4.0/" xlink:type="simple">
          <license-p>This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.</license-p>
        </license>
      </permissions>
      <abstract>
        <label>Abstract</label>
        <p>Coastal dune systems are dynamic and vulnerable landforms that provide key ecological, geomorphological and protective functions along the Bulgarian Black Sea coast. Despite their importance, their conservation status remains insufficiently quantified, particularly under the microtidal and fetch-limited conditions characteristic of the Black Sea.</p>
        <p>This study introduces the Eco-Geomorphological Dune Value Index (EGDVI), a multi-criteria assessment framework integrating 31 indicators across eight thematic categories, encompassing geomorphological, ecological, anthropogenic, legal and scientific dimensions. Unlike existing dune assessment approaches, the EGDVI combines these components within a single weighted and standardised scoring system, enabling a comprehensive and policy-relevant evaluation of dune systems. Indicator scores are derived from field surveys, high-resolution remote sensing data, GIS analysis and legislative sources and are weighted through structured expert elicitation.</p>
        <p>The EGDVI was applied to 16 dune systems along the Strandzha sector of the Bulgarian Black Sea coast. Total scores range from 154.0 to 364.5 (maximum 400), defining five classes of eco-geomorphological value. Four systems (25.0%) are classified as Very High Value (Class A), one (6.25%) as High Value (Class B), eight (50.0%) as Moderate Value (Class C), one (6.25%) as Low Value (Class D) and two (12.5%) as Degraded (Class E). At the regional scale, the mean EGDVI score (266.1) falls within the moderate-value class, indicating that transitional dune systems dominate along the Strandzha coast.</p>
        <p>The results demonstrate that the EGDVI provides a robust, transparent and reproducible framework for assessing both condition and conservation value. By explicitly incorporating governance and scientific dimensions alongside eco-geomorphological parameters, the index strengthens the science–policy interface and supports conservation prioritisation. The conceptual structure of the EGDVI is transferable to other microtidal and enclosed-sea coastal environments, provided that indicators are locally calibrated.</p>
      </abstract>
      <kwd-group>
        <label>Keywords</label>
        <kwd>anthropogenic pressure</kwd>
        <kwd>Bulgarian Black Sea Coast</kwd>
        <kwd>coastal dunes</kwd>
        <kwd>conservation assessment</kwd>
        <kwd>Eco-Geomorphological Dune Value Index</kwd>
      </kwd-group>
      <counts>
        <fig-count count="5"/>
        <table-count count="5"/>
        <ref-count count="107"/>
      </counts>
    </article-meta>
    <notes>
      <sec sec-type="Funding program">
        <title>Funding program</title>
        <p>Project “<italic>Mapping and Spatiotemporal Analysis of Beach-Dune Systems on the Southern Bulgarian Black Sea Coast: Evolution, Anthropogenic Pressure and Ecological Risks to Dune Habitats (MapBGBeachDune)</italic>”. Grant № КП-06-H84/5-16.12.2024 under Competition for financial support of basic research projects of the Bulgarian National Science Fund.</p>
      </sec>
    </notes>
  </front>
  <body>
    <sec sec-type="Introduction">
      <title>Introduction</title>
      <p>Coastal dune systems are formed by the interaction of marine, aeolian and terrestrial processes and are widely recognised as priority habitats in international conservation frameworks due to their ecological, geomorphological and socio-economic importance (<xref ref-type="bibr" rid="B13882570">Goldsmith 1985</xref>, <xref ref-type="bibr" rid="B13882830">Martínez et al. 2008</xref>, <xref ref-type="bibr" rid="B13882821">Martínez et al. 2013</xref>, <xref ref-type="bibr" rid="B13883059">Psuty and Silveira 2010</xref>, <xref ref-type="bibr" rid="B13882659">Hesp 2011</xref>, <xref ref-type="bibr" rid="B13882668">Hesp 2024</xref>, <xref ref-type="bibr" rid="B13882677">Hesp and Walker 2013</xref>, <xref ref-type="bibr" rid="B13882938">Nordstrom 2015</xref>, <xref ref-type="bibr" rid="B13882766">Luijendijk et al. 2018</xref>, <xref ref-type="bibr" rid="B13882340">Chadwick et al. 2022</xref>, <xref ref-type="bibr" rid="B13883036">Prodanov et al. 2023</xref>, <xref ref-type="bibr" rid="B13883104">Rangel-Buitrago et al. 2023</xref>, <xref ref-type="bibr" rid="B13882435">De Freitas 2024</xref>). Often described as part of the Coastal Transition Zone (CTZ), dune systems function as natural buffers that mitigate marine forcing, reduce shoreline erosion and preserve the integrity of the land–sea interface (<xref ref-type="bibr" rid="B13882839">Maun 2009</xref>). Beyond their protective role, dunes contribute to groundwater retention and purification, the regulation of local microclimates and the maintenance of habitats for highly specialised flora and fauna adapted to sandy, saline and wind-exposed environments (<xref ref-type="bibr" rid="B13882974">Pérez-Maqueo et al. 2013</xref>). As such, they deliver a broad range of ecosystem services, including storm and erosion protection, sediment storage for beach nourishment, carbon sequestration in dune vegetation and the provision of habitats for rare, endemic and threatened species (<xref ref-type="bibr" rid="B13882327">Carter et al. 1991</xref>, <xref ref-type="bibr" rid="B13882514">Feagin et al. 2005</xref>, <xref ref-type="bibr" rid="B13882482">Doody 2013</xref>). In addition, coastal dunes support recreational and tourism activities and hold cultural and educational value for coastal communities (<xref ref-type="bibr" rid="B13882929">Nordstrom 2000</xref>, <xref ref-type="bibr" rid="B13882974">Pérez-Maqueo et al. 2013</xref>, <xref ref-type="bibr" rid="B13882865">McLachlan et al. 2013</xref>). As integral components of sandy beach–dune systems, coastal dunes also function as dynamic land–sea interface units that ensure sediment continuity, ecological connectivity and natural buffering against marine forcing (<xref ref-type="bibr" rid="B13882857">McLachlan and Brown 2006</xref>).</p>
      <p>Coastal zones concentrate a wide range of natural, cultural and economic assets within a limited spatial extent, making them amongst the most intensively used and contested environments worldwide. Residential and recreational settlements, infrastructure, economic activities, specialised tourism (e.g. dunes as geomorphosites for geotourism), archaeological heritage and biodiversity-rich ecosystems are frequently juxtaposed within the same coastal space. Within this complex mosaic, beach-dune systems occupy a particularly sensitive position, as they must simultaneously sustain ecological functions, provide socio-economic services and absorb multiple forms of human pressure, especially in the context of sun-and-sea tourism (<xref ref-type="bibr" rid="B13882821">Martínez et al. 2013</xref>, <xref ref-type="bibr" rid="B13882812">Marrero-Rodríguez and Dóniz-Páez 2022</xref>, <xref ref-type="bibr" rid="B13882736">Johnston et al. 2023</xref>).</p>
      <p>Situated at the land–sea interface, coastal dune landforms are amongst the most dynamic geomorphic features, continuously reshaped by waves, storms and sea-level fluctuations, while also being strongly modified by human activities (<xref ref-type="bibr" rid="B13882719">Huggett 2011</xref>, <xref ref-type="bibr" rid="B13882874">Milne et al. 2012</xref>, <xref ref-type="bibr" rid="B13882532">Foti et al. 2021</xref>, <xref ref-type="bibr" rid="B13882417">D’Alessandro et al. 2022</xref>, <xref ref-type="bibr" rid="B13882523">Fontán-Bouzas et al. 2022</xref>, <xref ref-type="bibr" rid="B13883036">Prodanov et al. 2023</xref>, <xref ref-type="bibr" rid="B13883104">Rangel-Buitrago et al. 2023</xref>, <xref ref-type="bibr" rid="B13883264">Woodroffe et al. 2023</xref>, <xref ref-type="bibr" rid="B13883046">Prodanov et al. 2025</xref>). Their formation and evolution depend on three fundamental factors, sediment availability, wind regime and vegetation cover, resulting in high spatial and temporal variability shaped by the interplay of physical, chemical and biological processes (<xref ref-type="bibr" rid="B13883077">Pye 1983</xref>, <xref ref-type="bibr" rid="B13882514">Feagin et al. 2005</xref>, <xref ref-type="bibr" rid="B13883375">Aagaard et al. 2007</xref>, <xref ref-type="bibr" rid="B13883059">Psuty and Silveira 2010</xref>, <xref ref-type="bibr" rid="B13882241">Anthony 2013</xref>, <xref ref-type="bibr" rid="B13883420">Masselink et al. 2014</xref>, <xref ref-type="bibr" rid="B13882938">Nordstrom 2015</xref>, <xref ref-type="bibr" rid="B13883392">Delgado-Fernandez et al. 2019</xref>, <xref ref-type="bibr" rid="B13882499">Feagin et al. 2019</xref>, <xref ref-type="bibr" rid="B13883410">Costas et al. 2020</xref>).</p>
      <p>Despite their ecological, protective and socio-economic importance, coastal dunes are amongst the most threatened ecosystems worldwide (<xref ref-type="bibr" rid="B13882443">Defeo et al. 2009</xref>). Their conservation status reflects a delicate balance between natural drivers, such as sediment budgets, beach morphodynamics and vegetation succession and multiple anthropogenic pressures. Major human-induced threats include resource exploitation, urbanisation, coastal infrastructure development, unregulated tourism, trampling, off-road vehicle use, mechanical beach cleaning, invasive species and littering (<xref ref-type="bibr" rid="B13882650">Heslenfeld et al. 2008</xref>, <xref ref-type="bibr" rid="B13882318">Carboni et al. 2009</xref>, <xref ref-type="bibr" rid="B13882659">Hesp 2011</xref>, <xref ref-type="bibr" rid="B13882677">Hesp and Walker 2013</xref>, <xref ref-type="bibr" rid="B13882865">McLachlan et al. 2013</xref>, <xref ref-type="bibr" rid="B13882367">Ciccarelli 2014</xref>, <xref ref-type="bibr" rid="B13883246">Valcheva et al. 2018</xref>, <xref ref-type="bibr" rid="B13883255">Valcheva et al. 2021</xref>, <xref ref-type="bibr" rid="B13882883">Mo et al. 2021</xref>, <xref ref-type="bibr" rid="B13882376">Ciccarelli et al. 2023</xref>, <xref ref-type="bibr" rid="B13882668">Hesp 2024</xref>). These pressures are increasingly compounded by climate-related hazards, including accelerated sea-level rise, increased storm intensity and altered sediment supply, which drive erosion, dune mobility, habitat loss and reduced system resilience (<xref ref-type="bibr" rid="B13883122">Sabatier et al. 2009</xref>, <xref ref-type="bibr" rid="B13883086">Ranasinghe 2016</xref>, <xref ref-type="bibr" rid="B13882635">Harley et al. 2017</xref>, <xref ref-type="bibr" rid="B13882561">Gao et al. 2020</xref>). As a result, degradation of dune systems compromises their capacity to deliver ecosystem services, with direct implications for both ecological integrity and human well-being (<xref ref-type="bibr" rid="B13882617">Hanley et al. 2014</xref>, <xref ref-type="bibr" rid="B13882223">Alessandrini et al. 2024</xref>).</p>
      <p>The complexity and vulnerability of dune systems, coupled with their importance for ecosystem services, have spurred the development of targeted management and conservation strategies worldwide. Effective dune management requires integrated approaches that account for marine, aeolian and geological dynamics, while recognising the central role of vegetation in dune formation and stabilisation (<xref ref-type="bibr" rid="B13882839">Maun 2009</xref>). The overarching challenge lies in balancing habitat preservation with sustainable use by integrating ecological and physical processes with socio-economic and cultural considerations (<xref ref-type="bibr" rid="B13882232">Anfuso et al. 2014</xref>, <xref ref-type="bibr" rid="B13882912">Niavis and Tsiotas 2019</xref>, <xref ref-type="bibr" rid="B13882892">Mooser et al. 2022</xref>, <xref ref-type="bibr" rid="B13882490">Er-Ramy et al. 2022</xref>, <xref ref-type="bibr" rid="B13883095">Rangel-Buitrago and Gracia 2024</xref>). In this context, spatial mapping and classification emerge as fundamental management tools, forming the basis for coastal risk assessment and prioritisation of interventions. Coastal risk, commonly defined as the interaction between hazard probability and potential consequences, depends on the interplay between hazards and system vulnerability (<xref ref-type="bibr" rid="B13882686">Hieronymus and Kalén 2020</xref>). Existing coastal risk and vulnerability indices often prioritise physical and geological parameters, while incorporating socio-economic, ecological and cultural aspects to a more limited extent (<xref ref-type="bibr" rid="B13882903">Nguyen et al. 2016</xref>, <xref ref-type="bibr" rid="B13883113">Rocha et al. 2023</xref>). Management responses range from passive measures such as land-use zoning and legal protection to active interventions including dune stabilisation, habitat restoration and invasive species control (<xref ref-type="bibr" rid="B13882821">Martínez et al. 2013</xref>, <xref ref-type="bibr" rid="B13882938">Nordstrom 2015</xref>). However, these measures require robust, standardised assessment frameworks capable of integrating physical, ecological and socio-economic information to support evidence-based decision-making (<xref ref-type="bibr" rid="B13882552">Gallego-Fernández et al. 2011</xref>, <xref ref-type="bibr" rid="B13882250">Arens et al. 2013</xref>, <xref ref-type="bibr" rid="B13882541">Furlan et al. 2021</xref>). However, human interventions frequently constrain natural dune dynamics by altering sediment pathways, vegetation succession and morphological feedbacks, underscoring the need for assessment frameworks that explicitly account for anthropogenic pressure and management intensity (<xref ref-type="bibr" rid="B13882947">Nordstrom et al. 2015</xref>).</p>
      <p>Recent coastal assessment frameworks increasingly emphasise the integration of geomorphological integrity, environmental quality, anthropogenic disturbance, and ecosystem functionality within unified management-oriented approaches (<xref ref-type="bibr" rid="B13882757">Lithgow et al. 2015</xref>, <xref ref-type="bibr" rid="B13882994">Pinna et al. 2022</xref>, <xref ref-type="bibr" rid="B13882456">Della Bella et al. 2024</xref>, <xref ref-type="bibr" rid="B14222549">Rangel-Buitrago et al. 2026</xref>). Numerous indices and assessment tools have been developed to evaluate dune condition, vulnerability and conservation status, reflecting different disciplinary perspectives and management objectives. These approaches can be broadly grouped into four categories. Ecological and vegetation-based indices focus on floristic diversity, vegetation structure and habitat quality as indicators of dune condition, including the H′dune index (<xref ref-type="bibr" rid="B13882579">Grunewald and Schubert 2007</xref>), the Mediterranean conservation status framework proposed by <xref ref-type="bibr" rid="B13882318">Carboni et al. (2009)</xref> and the vegetation-based risk index developed by <xref ref-type="bibr" rid="B13882223">Alessandrini et al. (2024)</xref>. Geomorphological and physical condition indices emphasise dune morphology, sediment dynamics and exposure to erosive forces, including restoration-orientated approaches such as those discussed by <xref ref-type="bibr" rid="B13882757">Lithgow et al. (2015)</xref> and regional vulnerability frameworks (<xref ref-type="bibr" rid="B13882964">Peña-Alonso et al. 2018</xref>), as well as vegetation-based conservation indices applied by <xref ref-type="bibr" rid="B13883003">Pinna et al. (2015)</xref>. Integrated eco-geomorphological indices combine physical and biological parameters to capture system complexity, notably the Coastal Dune Vulnerability Index (CDVI) and its subsequent refinements (<xref ref-type="bibr" rid="B13882388">Ciccarelli et al. 2017</xref>, <xref ref-type="bibr" rid="B13882287">Bertoni et al. 2019</xref>), as well as endemicity-focused extensions (<xref ref-type="bibr" rid="B13882985">Pinna et al. 2019</xref>). Finally, multidimensional socio-ecological and legal frameworks incorporate a broad set of drivers, including climate forcing, human use and legal protection, as exemplified by the Multidimensional Coastal Vulnerability Index (<xref ref-type="bibr" rid="B13882541">Furlan et al. 2021</xref>) and by checklist-based approaches (<xref ref-type="bibr" rid="B13886304">Bodéré et al. 1991</xref>, <xref ref-type="bibr" rid="B13882296">Bodéré et al. 1994</xref>), alongside restoration-focused tools such as the Dune Restoration Potential index (<xref ref-type="bibr" rid="B13882994">Pinna et al. 2022</xref>).</p>
      <p>Despite their diversity, existing frameworks share several common limitations. Many are strongly context-dependent, hindering transferability without extensive local recalibration (<xref ref-type="bibr" rid="B13883068">Pye and Blott 2008</xref>, <xref ref-type="bibr" rid="B13882552">Gallego-Fernández et al. 2011</xref>). Others tend to over-emphasise either physical or ecological dimensions while under-representing complementary aspects. In particular, legal protection status, scientific research value and nuanced forms of anthropogenic pressure are rarely quantified in a systematic manner, limiting the usefulness of many indices for policy-orientated conservation planning (<xref ref-type="bibr" rid="B13882541">Furlan et al. 2021</xref>, <xref ref-type="bibr" rid="B13882223">Alessandrini et al. 2024</xref>). Differences in indicator selection, weighting schemes and scoring approaches further reduce comparability and repeatability across studies.</p>
      <p>Addressing these gaps, the present study introduces the EGDVI. This unified, standardised and transferable framework integrates geomorphology, ecology, human pressures, legal protection and scientific value into a single operational assessment tool. The EGDVI is specifically adapted to the microtidal, fetch-limited conditions of the Black Sea coast, where dune dynamics, management practices and conservation challenges differ markedly from those of macro- and mesotidal environments.</p>
      <sec sec-type="Aim of the study">
        <title>Aim of the study</title>
        <p>This study aims to develop and apply the EGDVI, a multi-criteria evaluation framework designed to overcome the limitations of existing dune assessment methodologies. The EGDVI comprises 31 indicators grouped into eight categories: <italic>Geomorphology</italic> (GM), <italic>Aeolian Dynamics</italic> (AD), <italic>Ecology</italic> (ECO), <italic>Pressure</italic> (PRS), <italic>Human Footprint</italic> (HFP), <italic>Legal Protection</italic> (LEG), <italic>Scientific Value</italic> (SCI) and <italic>Nature Conservation</italic> (CON). All indicators are scored on a standardised 0–4 scale, where higher scores indicate higher natural value or better conservation condition and are weighted, based on expert assessment and scientific review.</p>
        <p>The framework is applied to the Strandzha dune systems along the southern Bulgarian Black Sea Coast, a region characterised by diverse coastal morphologies ranging from narrow pocket beaches backed by cliffs to broad, low-lying barrier systems. Although the Black Sea is microtidal, it exhibits complex sediment dynamics and distinct erosion behaviour associated with wind-driven waves and storm surges (<xref ref-type="bibr" rid="B13883156">Tătui et al. 2019</xref>). At the same time, most beaches are concession-managed and subject to regular levelling and mechanical cleaning, which obscures their natural state. For this reason, dune systems were selected as the primary units of assessment. This context enables the inclusion of Black Sea–specific indicators, enhancing the regional relevance of the EGDVI, while demonstrating its potential transferability to other low-energy coastal environments.</p>
      </sec>
    </sec>
    <sec sec-type="Materials and methods">
      <title>Materials and methods</title>
      <sec sec-type="Pilot Assessment Dune Systems">
        <title>Pilot Assessment Dune Systems</title>
        <sec sec-type="Dune systems along the Strandzha Coast (southern Bulgarian Black Sea Coast)">
          <title>Dune systems along the Strandzha Coast (southern Bulgarian Black Sea Coast)</title>
          <p>The Strandzha sector of the southern Bulgarian Black Sea Coast hosts a diverse set of coastal dune systems shaped by the interaction of aeolian, marine and fluvial processes within a geologically complex setting of alternating rocky headlands, pocket beaches and sediment-rich embayments. Sixteen distinct beach–dune systems have been identified, covering 101.89 ha, or approximately 10.3% of Bulgaria’s dune area. Primorsko (International Youth Centre) is the largest at 26.73 ha (<xref ref-type="bibr" rid="B13883012">Prodanov et al. 2026</xref>). Human pressure is a major driver of change: 4.8 ha of dunes are degraded due to tourism, infrastructure, beach maintenance and unregulated access. The most affected dune systems include Ahtopol, Kiten, Koral and Stomopolu–Perla (<xref ref-type="bibr" rid="B13883036">Prodanov et al. 2023</xref>, <xref ref-type="bibr" rid="B13883012">Prodanov et al. 2026</xref>).</p>
          <p>Morphologically, these systems include incipient (embryonic) foredunes, established foredunes, river-influenced foredunes, blowouts, parabolic dunes (including climbing and cliff-top forms) and transgressive dunefields (<xref ref-type="bibr" rid="B13883036">Prodanov et al. 2023</xref>, <xref ref-type="bibr" rid="B13883046">Prodanov et al. 2025</xref>, <xref ref-type="bibr" rid="B13883012">Prodanov et al. 2026</xref>) – (Figs <xref ref-type="fig" rid="F13883313">1</xref>, <xref ref-type="fig" rid="F13883315">2</xref>, <xref ref-type="fig" rid="F13883317">3</xref>). Each type reflects specific morphodynamic conditions, sediment supply regimes and vegetation–wind interactions.</p>
          <p>Incipient foredunes occur across most systems, often as nebkhas or ramps and are highly sensitive to human disturbance. Established foredunes, some exceeding 18 m in height and 150 m in width, form prominent protective ridges. River-influenced foredunes, notably at the mouth of the Veleka River, exhibit periodic erosion and recovery linked to flood events. Blowouts and parabolic dunes illustrate the balance between wind-driven erosion and vegetation-led stabilisation. At the same time, transgressive dunefields, such as those at Primorsko (International Youth Centre), represent the most dynamic environments (<xref ref-type="bibr" rid="B13883012">Prodanov et al. 2026</xref>).</p>
          <p>The EUNIS habitat classification (2020) identifies sand dunes from the Bulgarian Black Sea Coast as: N12 Mediterranean and Black Sea sand beach; N14 Mediterranean, Macronesian and Black Sea shifting coastal dune; N17 Black Sea coastal dune grassland (grey dune); N1E Black Sea broad-leaved coastal dune forest; N1J Mediterranean and Black Sea moist and wet dune slack. These habitat types respond to the following Natura 2000 habitats: 2110 *Embryonic shifting dunes; 2120 *Shifting dunes along the shoreline with <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Ammophila">Ammophila</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="arenaria">arenaria</tp:taxon-name-part></tp:taxon-name></italic> (white dunes); 2130 *Fixed coastal dunes with herbaceous vegetation (grey dunes); 2180 *Wooded dunes of the Atlantic, Continental and Boreal region; 2190 *Humid dune slacks.</p>
          <p>Psammophytic vegetation from the Bulgarian Black Sea Coast includes embryonic dunes of <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Cakiletea">Cakiletea</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="maritimae">maritimae</tp:taxon-name-part></tp:taxon-name></italic>, represented by one alliance (<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Euphorbion">Euphorbion</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="peplidis">peplidis</tp:taxon-name-part></tp:taxon-name></italic>) with one association (<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Cakilo">Cakilo</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="euxinae">euxinae</tp:taxon-name-part></tp:taxon-name></italic>–<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Salsoletum">Salsoletum</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="ruthenicae">ruthenicae</tp:taxon-name-part></tp:taxon-name></italic>) and white and grey dunes of class <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Ammophiletea">Ammophiletea</tp:taxon-name-part></tp:taxon-name></italic>, with three alliances (<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Elymion">Elymion</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="gigantei">gigantei</tp:taxon-name-part></tp:taxon-name></italic> with two associations, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Xanthio">Xanthio</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="italici">italici</tp:taxon-name-part></tp:taxon-name></italic>–<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Leymetum">Leymetum</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="sabulosi">sabulosi</tp:taxon-name-part></tp:taxon-name></italic> and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Medicago">Medicago</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="tenderiensis">tenderiensis</tp:taxon-name-part></tp:taxon-name></italic>–<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Ammophiletum">Ammophiletum</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="arundinaceae">arundinaceae</tp:taxon-name-part></tp:taxon-name></italic>; <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Scabiosion">Scabiosion</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="ucrainicae">ucrainicae</tp:taxon-name-part></tp:taxon-name></italic> with three associations, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Alysso">Alysso</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="borzaeani">borzaeani</tp:taxon-name-part></tp:taxon-name></italic>–<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Ephedretum">Ephedretum</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="distachyae">distachyae</tp:taxon-name-part></tp:taxon-name></italic>, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Stachyo">Stachyo</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="atherocalici">atherocalici</tp:taxon-name-part></tp:taxon-name></italic>–<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Caricetum">Caricetum</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="ligericae">ligericae</tp:taxon-name-part></tp:taxon-name></italic> and <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Festuca">Festuca</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="arenicola">arenicola</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="subspecies" reg="community">community</tp:taxon-name-part></tp:taxon-name></italic>; <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Sileno">Sileno</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="thymifolae">thymifolae</tp:taxon-name-part></tp:taxon-name></italic>–<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Jurinion">Jurinion</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="kilaeae">kilaeae</tp:taxon-name-part></tp:taxon-name></italic> with one association, <italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Aurinio">Aurinio</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="uechtritziani">uechtritziani</tp:taxon-name-part></tp:taxon-name></italic>–<italic><tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Artemisietum">Artemisietum</tp:taxon-name-part> <tp:taxon-name-part taxon-name-part-type="species" reg="campestris">campestris</tp:taxon-name-part></tp:taxon-name></italic>) (<xref ref-type="bibr" rid="B13883220">Tzonev et al. 2005</xref>, <xref ref-type="bibr" rid="B13882787">Marcenò et al. 2018</xref>).</p>
          <p>In addition to their geomorphological and ecological diversity, the Strandzha dune systems are embedded in a strict legal and institutional framework. According to Article 18 of the Constitution of the Republic of Bulgaria (<xref ref-type="bibr" rid="B13882401">CRB 1991</xref>), the coastal beach strip, including dunes, is exclusive state property. Their protection is further regulated by the Black Sea Coast Spatial Development Act (<xref ref-type="bibr" rid="B13882310">BSCSDA 2007</xref>), which provides the first national legal definition of sand dunes and requires the preparation of specialised cadastral maps and registers (<xref ref-type="bibr" rid="B13882956">Ordinance 2008</xref>). Complementary provisions in the Biological Diversity Act (<xref ref-type="bibr" rid="B13882279">BDA 2002</xref>) ensure that dune habitats are recognised as natural habitats of conservation importance, harmonised with Annex I of the EU Habitats Directive (<xref ref-type="bibr" rid="B13882409">Council Directive 92/43/EEC 1992</xref>). Consequently, all five dune habitat types present in Strandzha are formally protected under Bulgarian and EU law (<xref ref-type="bibr" rid="B13882279">BDA 2002</xref>, <xref ref-type="bibr" rid="B13882609">Gussev and Tzonev 2014</xref>, <xref ref-type="bibr" rid="B13882588">Gussev 2015</xref>, <xref ref-type="bibr" rid="B13883175">Tzonev 2015a</xref>, <xref ref-type="bibr" rid="B13883184">Tzonev 2015b</xref>, <xref ref-type="bibr" rid="B13883193">Tzonev 2015c</xref>, <xref ref-type="bibr" rid="B13883202">Tzonev 2015d</xref>, <xref ref-type="bibr" rid="B13883211">Tzonev 2015e</xref>, <xref ref-type="bibr" rid="B13883229">Tzonev and Gussev 2015</xref>, <xref ref-type="bibr" rid="B13883237">Valcheva et al. 2020</xref>, <xref ref-type="bibr" rid="B13883255">Valcheva et al. 2021</xref>).</p>
          <p>Despite this legal recognition, enforcement has been inconsistent. The national cadastral system (CAIS) suffers from outdated or incomplete dune mapping, which has led to degradation, land-use conflicts and unauthorised construction on dunes at several Strandzha sites (<xref ref-type="bibr" rid="B13883036">Prodanov et al. 2023</xref>, <xref ref-type="bibr" rid="B13883027">Prodanov and Bekova 2024</xref>). To address these gaps, the Ministry of Environment and Water, together with the Institute of Oceanology ant the Institute of Biodiversity and Ecosystem Research at the Bulgarian Academy of Sciences, developed the Standard Dune Mapping Procedure (SDMP), which mandates high-resolution UAV photogrammetry, geomorphological and phytocoenological surveys and the production of authoritative cadastral maps at 1:1000 scale (<xref ref-type="bibr" rid="B13883046">Prodanov et al. 2025</xref>).</p>
          <p>Thus, the Strandzha Black Sea Coast is not only a hotspot of geomorphological and ecological richness, but also a test ground for the implementation of Bulgaria’s dune protection laws and new standardised mapping methodologies. The combination of high natural value and strict, yet imperfectly enforced, legal safeguards makes it an ideal pilot area for applying the EGDVI to bridge science, conservation and policy.</p>
        </sec>
      </sec>
      <sec sec-type="Methods for the development of EGDVI">
        <title>Methods for the development of EGDVI</title>
        <p>The proposed EGDVI is a multi-criteria assessment framework that integrates geomorphological, aeolian, ecological, anthropogenic, legal, scientific and conservation aspects into a single, transferable methodology for evaluating the natural value and condition of dune systems. The index comprises 31 indicators grouped into eight thematic categories, each reflecting a distinct dimension of eco-geomorphological value. They include <italic>Geomorphology (GM)</italic> – 5 indicators; <italic>Aeolian Dynamics (AD)</italic> – 4 indicators, <italic>Ecology (ECO)</italic> – 5 indicators, <italic>Pressure/Risk/Stress (PRS)</italic> – 4 indicators, <italic>Human Footprint Pressure (HFP)</italic> – 4 indicators, <italic>Legal/Protection Status/Management (LEG)</italic> – 5 indicators, <italic>Scientific Importance (SCI)</italic> – 2 indicators, <italic>Nature Conservation (CON)</italic> – 2 indicators. Indicators were selected, based on an extensive review of scientific literature, expert assessment and the specific characteristics of the Bulgarian Black Sea coast (Table <xref ref-type="table" rid="T13883321">1</xref>; Suppl. material <xref ref-type="supplementary-material" rid="S13883629">1</xref>).</p>
        <sec sec-type="Selection of indicators">
          <title>Selection of indicators</title>
          <p>Indicators were selected through:</p>
          <p><list list-type="bullet">
            <list-item>
              <p>A comprehensive review of existing dune evaluation methodologies (ecological, geomorphological, legal and socio-ecological frameworks) (<xref ref-type="bibr" rid="B13886304">Bodéré et al. 1991</xref>, <xref ref-type="bibr" rid="B13886318">García-Mora et al. 2001</xref>, <xref ref-type="bibr" rid="B13882579">Grunewald and Schubert 2007</xref>, <xref ref-type="bibr" rid="B13882318">Carboni et al. 2009</xref>, <xref ref-type="bibr" rid="B13882541">Furlan et al. 2021</xref>, <xref ref-type="bibr" rid="B13883003">Pinna et al. 2015</xref>, <xref ref-type="bibr" rid="B13882388">Ciccarelli et al. 2017</xref>, <xref ref-type="bibr" rid="B13882287">Bertoni et al. 2019</xref>, <xref ref-type="bibr" rid="B13882994">Pinna et al. 2022</xref>, <xref ref-type="bibr" rid="B13882223">Alessandrini et al. 2024</xref>, <xref ref-type="bibr" rid="B13882456">Della Bella et al. 2024</xref>);</p>
            </list-item>
            <list-item>
              <p>Adaptation to the specific conditions of the microtidal, fetch-limited Black Sea Coast, where dune systems are the most reliable natural units for conservation assessment;</p>
            </list-item>
            <list-item>
              <p>Consultation with domain experts in coastal geomorphology, dune ecology and psammophytic vegetation and environmental management;</p>
            </list-item>
            <list-item>
              <p>Field validation during May – September 2025 – pilot surveys along the Strandzha Black Sea Coast.</p>
            </list-item>
          </list></p>
          <p>Each indicator is scored on a standardised 0–4 scale, where 0 denotes the absence of value or the maximum risk/pressure and 4 reflects the maximum eco-geomorphological value or the optimal state. For pressure-type indicators (e.g. degradation risk), the scoring logic is inverted so that higher scores correspond to lower risk, ensuring consistency across all categories.</p>
        </sec>
        <sec sec-type="Indicator weighting and expert elicitation">
          <title>Indicator weighting and expert elicitation</title>
          <p>Indicator weights were derived through a structured expert-elicitation process to ensure transparency and repeatability. An interdisciplinary panel of coastal geomorphologists, dune ecologists and conservation practitioners first submitted independent weight proposals, each accompanied by justifications for each indicator and category. A second consensus round then reconciled differences and normalised the final weights to sum to 100%. This approach balances a literature-based rationale with documented expert consensus, while avoiding ad hoc weighting. Category totals are: GM 30%, AD 6%, ECO 26%, PRS 7.5%, HFP 9.5%, LEG 10%, SCI 4%, CON 7% (Σ = 100%) – Table <xref ref-type="table" rid="T13883321">1</xref>; Suppl. material <xref ref-type="supplementary-material" rid="S13886327">2</xref>; Suppl. material <xref ref-type="supplementary-material" rid="S13886328">3</xref>. Indicator weights were scaled proportionally within each category to reflect diagnostic power, measurability and management relevance. Where data for a given indicator were unavailable, its weight was redistributed within the same category to preserve the category total.</p>
        </sec>
        <sec sec-type="Calculating EGDVI">
          <title>Calculating EGDVI</title>
          <p>All indicator scoring, weighting procedures and final EGDVI calculations were implemented using an Excel-based assessment tool developed specifically for the application of the EGDVI to the Strandzha dune systems (Suppl. material <xref ref-type="supplementary-material" rid="S13886329">4</xref>). The tool implements the calculation workflow described below, ensuring consistent application of weights, automated score aggregation and class assignment.</p>
          <p>The <bold>Weighted Score (<italic>WS</italic>)</bold> for each indicator is calculated as:</p>
          <p>
            <tex-math id="M1">\documentclass[12pt]{standalone}
\usepackage{varwidth}

\usepackage[utf8x]{inputenc}
\usepackage[T1]{fontenc}
\usepackage{lmodern}

\usepackage{amsmath, amssymb, graphics, setspace}
\newcommand{\mathsym}[1]{{}}
\newcommand{\unicode}[1]{{}}
\newcounter{mathematicapage}
\begin{document}
   \begin{varwidth}{50in}
        \begin{equation*}
            WS_i = S_i \times W_i
        \end{equation*}
    \end{varwidth}
\end{document}
</tex-math>
          </p>
          <p>where <italic><bold>Si</bold></italic> is the indicator score (0–4) and <italic><bold>Wi</bold></italic> is the indicator weight expressed as a percentage, with the sum of all Wi equal to 100.</p>
          <p>The total EGDVI score for a given dune system is calculated as:</p>
          <p><tex-math id="M2">\documentclass[12pt]{standalone}
\usepackage{varwidth}

\usepackage[utf8x]{inputenc}
\usepackage[T1]{fontenc}
\usepackage{lmodern}

\usepackage{amsmath, amssymb, graphics, setspace}
\newcommand{\mathsym}[1]{{}}
\newcommand{\unicode}[1]{{}}
\newcounter{mathematicapage}
\begin{document}
   \begin{varwidth}{50in}
        \begin{equation*}
            EGDVI_{Total} = \sum_{i=1}^{n} (S_i \times W_i)
        \end{equation*}
    \end{varwidth}
\end{document}
</tex-math>,</p>
          <p>where n is the total number of indicators used in the index (n = 31 in the present study).</p>
          <p>As each indicator score (<italic><bold>Si</bold></italic>) can reach a maximum value of 4 and the sum of all indicator weights equals 100, the theoretical maximum EGDVI score is:</p>
          <p><tex-math id="M3">\documentclass[12pt]{standalone}
\usepackage{varwidth}

\usepackage[utf8x]{inputenc}
\usepackage[T1]{fontenc}
\usepackage{lmodern}

\usepackage{amsmath, amssymb, graphics, setspace}
\newcommand{\mathsym}[1]{{}}
\newcommand{\unicode}[1]{{}}
\newcounter{mathematicapage}
\begin{document}
   \begin{varwidth}{50in}
        \begin{equation*}
            EGDVI_{\mathrm{max}} = 4 \times 100 = 400
        \end{equation*}
    \end{varwidth}
\end{document}
</tex-math>,</p>
          <p>which corresponds to an ideal state in which all indicators reach their highest possible values.</p>
          <p>To improve the clarity of the calculation procedure, a brief illustrative example is provided. Consider a dune system evaluated using three representative indicators: GM-1 (Geomorphological Diversity), ECO-1 (Vegetation Integrity) and PRS-2 (Degradation Risk).</p>
          <p>Assuming scores of S = 4, S = 3 and S = 2 and corresponding weights of W = 6%, W = 5.5% and W = 2.5%, the weighted scores (WS), calculated as S × W, are 24.0, 16.5 and 5.0, respectively. The partial sum of these indicators is therefore 45.5.</p>
          <p>The final EGDVI score is obtained by summing the weighted scores of all indicators (n = 31), as implemented in the Excel-based assessment tool (Suppl. material <xref ref-type="supplementary-material" rid="S13886329">4</xref>). This example illustrates the additive and weighted structure of the index.</p>
        </sec>
        <sec sec-type="Classification of EGDVI scores">
          <title>Classification of EGDVI scores</title>
          <p>Classification into five conservation value classes followed a two-step approach:</p>
          <p><list list-type="order">
            <list-item>
              <p>Jenks Natural Breaks optimisation (<xref ref-type="bibr" rid="B13882727">Jenks 1967</xref>) was first applied to the pilot dataset (16 dune systems) to identify natural groupings and thresholds. This provided a data-driven basis for distinguishing distinct clusters and “<italic>knife-edge</italic>” cases;</p>
            </list-item>
            <list-item>
              <p>To ensure long-term comparability and policy applicability, we defined fixed reference thresholds as percentages of the maximum score (400): Class A – Very High: ≥ 320 (≥ 80%); Class B – High: 280–319 (70–79%); Class C – Moderate: 220–279 (55–69%); Class D – Low: 160–219 (40–54%); Class E – Degraded: ≤ 159 (≤ 39%) – Table <xref ref-type="table" rid="T13883322">2</xref>;</p>
            </list-item>
            <list-item>
              <p>Class assignment and threshold-based classification were performed automatically within the Excel-based assessment tool, following the criteria defined above (Suppl. material <xref ref-type="supplementary-material" rid="S13886329">4</xref>).</p>
            </list-item>
          </list></p>
          <p>This hybrid strategy preserves Jenks’s statistical robustness for initial diagnostics, while ensuring stable, transferable benchmarks for national monitoring frameworks and future assessments.</p>
        </sec>
        <sec sec-type="Data sources and measurement methods">
          <title>Data sources and measurement methods</title>
          <p>Indicator scoring was based on multiple data sources (Table <xref ref-type="table" rid="T13883321">1</xref>):</p>
          <p><list list-type="bullet">
            <list-item>
              <p>Field surveys: geomorphological mapping, GPS profiling, vegetation characteristics and structure and habitat mapping;</p>
            </list-item>
            <list-item>
              <p>Remote sensing: UAV photogrammetry, LiDAR transects and orthophoto interpretation for dune extent and morphology;</p>
            </list-item>
            <list-item>
              <p>GIS analysis: shoreline position, dune width, blowout extent and infrastructure footprint;</p>
            </list-item>
            <list-item>
              <p>Legislative/cadastral databases: land ownership, protection status and zoning;</p>
            </list-item>
            <list-item>
              <p>Scientific literature and inventories: habitat types, floristic and phytocoenological data and geodiversity records.</p>
            </list-item>
          </list></p>
          <p>When data for a given indicator were unavailable, its weight was redistributed proportionally within the same category to maintain the overall balance.</p>
        </sec>
        <sec sec-type="Sensitivity of EGDVI">
          <title>Sensitivity of EGDVI</title>
          <p>To evaluate the robustness of the EGDVI, we conducted sensitivity analyses to assess how uncertainties in indicator weighting affect the ranking and classification of dune systems. Two complementary approaches were used:</p>
          <p><list list-type="order">
            <list-item>
              <p><italic>One-at-a-time (OAT)</italic> sensitivity analysis, where individual indicator weights were varied by ± 10–20% of their baseline values (renormalised to 100%), while scores remained constant. This allowed us to test the local stability of site rankings under small perturbations;</p>
            </list-item>
            <list-item>
              <p><italic>Category-level perturbation scenarios</italic> (<xref ref-type="bibr" rid="B13882903">Nguyen et al. 2016</xref>), where weight shares were shifted by up to ± 15% between paired thematic categories (e.g., ECO ↔ GM, PRS ↔ HFP). This procedure explored broader redistributions of expert-derived weights, reflecting realistic uncertainty in expert assessment.</p>
            </list-item>
          </list></p>
          <p>Across both OAT and scenario-based perturbations, the rank order of dune systems remained highly stable, with changes limited to ≤ 1 class step and Spearman’s rank correlation between baseline and perturbed ranks exceeding 0.90. This demonstrates that the EGDVI is robust to plausible levels of weighting uncertainty and that its classification outcomes are reliable for conservation and policy applications.</p>
        </sec>
      </sec>
    </sec>
    <sec sec-type="Results">
      <title>Results</title>
      <p>Application of the Eco-Geomorphological Dune Value Index (EGDVI) to the 16 dune systems along the Strandzha coast reveals a pronounced differentiation in eco-geomorphological condition across the region. Total scores range from 154.0 to 364.5, out of a theoretical maximum of 400, indicating the co-existence of both highly preserved and severely degraded dune systems within a relatively compact coastal sector.</p>
      <p>Based on the fixed classification thresholds (Fig. <xref ref-type="fig" rid="F13883319">4</xref>; Tables <xref ref-type="table" rid="T13883325">3</xref>, <xref ref-type="table" rid="T13883326">4</xref>), the assessed dune systems are unevenly distributed across value classes. Very high eco-geomorphological value (Class A) characterises four systems (25.0%), while high value (Class B) is represented by a single system (6.25%). The majority of dune systems (eight sites, 50.0%) fall within Class C – Moderate Value, reflecting intermediate condition. Low-value (Class D) and degraded systems (Class E) together account for three sites (18.75%), underscoring the presence of localised, but significant degradation hotspots along the coast.</p>
      <p>Summary statistics (Table <xref ref-type="table" rid="T13883326">4</xref>) indicate a strongly asymmetric distribution of EGDVI scores. Although a limited number of dune systems attain very high values, the regional mean score (266.1) and median (261.2) both fall within the moderate-value range. This highlights the dominance of transitional systems at the scale of the Strandzha coast, rather than a landscape uniformly characterised by either high conservation status or widespread degradation.</p>
      <sec sec-type="Class A – Very High Eco-Geomorphological Value">
        <title>Class A – Very High Eco-Geomorphological Value</title>
        <p>The highest eco-geomorphological values are recorded for Primorsko (Stomopolu–Perla), Primorsko (International Youth Centre), Koral and Camping Yug and Lipite (Fig. <xref ref-type="fig" rid="F13883319">4</xref>). These four systems form a distinct upper cluster, with EGDVI scores consistently exceeding 320 and a high mean value (347.4). Their relatively narrow interquartile range (IQR = 19.4) indicates limited internal variability and suggests that all four systems approach near-reference conditions.</p>
        <p>From a geomorphological perspective, these dune systems are characterised by wide, continuous dune belts that often exceed 100–150 m and by well-developed morphological sequences, including incipient dunes, foredunes, secondary ridges and, locally, parabolic or cliff-top dune forms. Ecologically, they exhibit high vegetation integrity with clear zonation patterns, while anthropogenic pressure remains comparatively low. Favourable legal protection status and feasible conservation management further contribute to their consistently high scores.</p>
      </sec>
      <sec sec-type="Class B – High Eco-Geomorphological Value">
        <title>Class B – High Eco-Geomorphological Value</title>
        <p>Atliman represents the sole dune system classified as Class B – High Value. With an EGDVI score of 283.5, the system occupies a transitional position immediately below the Class A threshold. This indicates a high degree of geomorphological and ecological integrity, albeit constrained by localised pressures, such as regulated access and seasonal recreational use.</p>
        <p>Although statistical dispersion measures are not applicable to a single system, Atliman’s proximity to the upper-class boundary suggests that relatively minor changes in management effectiveness could result in upward or downward reclassification. As such, the system exemplifies the sensitivity of high-value dune environments to subtle shifts in anthropogenic pressure.</p>
      </sec>
      <sec sec-type="Class C – Moderate Eco-Geomorphological Value">
        <title>Class C – Moderate Eco-Geomorphological Value</title>
        <p>Class C – Moderate Value encompasses the largest and most regionally representative group, including Veleka River Mouth, Kiten, Butamyata, Ahtopol, Listi, Silistar, Nestinarka and Oazis (Fig. <xref ref-type="fig" rid="F13883319">4</xref>). With a mean score of 258.9 and a median of 259.2, these systems collectively define the prevailing eco-geomorphological condition along the Strandzha coast.</p>
        <p>Despite their numerical dominance, Class C systems display relatively limited internal dispersion (IQR = 7.6), indicating broadly comparable levels of condition. Most retain functional dune units and key geomorphological elements, yet exhibit varying degrees of fragmentation, reduced sediment connectivity and visible anthropogenic influence, particularly within the foredune zone. The presence of several systems close to the upper threshold of this class, such as the Veleka River Mouth, highlights the potential for targeted management interventions to yield substantial improvements.</p>
      </sec>
      <sec sec-type="Class D – Low Eco-Geomorphological Value">
        <title>Class D – Low Eco-Geomorphological Value</title>
        <p>Lozenets is classified as Class D – Low Value, reflecting a markedly reduced eco-geomorphological condition relative to the regional average. The system is characterised by a narrow, discontinuous dune belt, disrupted vegetation cover and diminished morphodynamic function. Although not yet fully degraded, its low score indicates limited resilience and a reduced capacity for natural recovery under current pressure regimes.</p>
      </sec>
      <sec sec-type="Class E – Degraded Dune Systems">
        <title>Class E – Degraded Dune Systems</title>
        <p>The lowest EGDVI scores are observed at Tsarevo (Popski plazh) and Arapya, both classified as Class E – Degraded. These systems exhibit consistently low values with minimal variability, reflecting a uniformly advanced state of degradation. Extensive loss of frontal dune ridges, sparse psammophytic vegetation and a near absence of coastal-protection function characterise both sites.</p>
        <p>Long-standing anthropogenic pressure, compounded by shoreline modification and storm impacts, has resulted in persistent degradation that is unlikely to be reversed without substantial and sustained restoration measures.</p>
      </sec>
      <sec sec-type="Regional patterns and management implications">
        <title>Regional patterns and management implications</title>
        <p>Comparison between class-level statistics and the overall distribution of EGDVI scores highlights pronounced spatial heterogeneity along the Strandzha coast (regional IQR = 44.5). While a small number of dune systems retain near-reference conditions, the prevalence of moderately valued systems underscores the importance of proactive management. These transitional systems represent both a risk if pressure continues unchecked and an opportunity, as relatively modest interventions could significantly enhance their eco-geomorphological value.</p>
        <p>Overall, the results demonstrate that the Strandzha coast simultaneously hosts some of the most valuable and some of the most degraded dune systems in Bulgaria, confirming the EGDVI’s effectiveness in discriminating between levels of condition and in supporting conservation prioritisation and evidence-based coastal management.</p>
      </sec>
      <sec sec-type="Case Study: Primorsko (Stomopolu–Perla) Dune System">
        <title>Case Study: Primorsko (Stomopolu–Perla) Dune System</title>
        <p>To better illustrate how the EGDVI performs in a real-world context, a representative dune system from the Strandzha coast, Primorsko (Stomopolu–Perla), is examined in detail. The site reaches a total score of 364.5, placing it in Class A (Very High Eco-Geomorphological Value), which corresponds to a near-pristine system with outstanding geomorphological structure, ecological integrity and conservation importance (Fig. <xref ref-type="fig" rid="F14194734">5</xref>).</p>
        <p>All geomorphological indicators (GM-1 to GM-5) receive the maximum score (4), pointing to a well-developed and continuous dune system with clear zonation from embryonic dunes to foredunes and secondary ridges. The dunes are both high (&gt; 5 m) and wide (&gt; 100 m), which places them in the highest class definitions. Hydrogeomorphic connectivity is fully functional, allowing active sediment and water exchange with adjacent environments. At the same time, the lack of fragmentation confirms that the system functions as a continuous, structurally intact dune corridor.</p>
        <p>Indicators related to aeolian dynamics (AD-1 to AD-4 = 3) suggest a system that is active, but stable. Blowouts are present, but limited in extent (&lt; 10%) and sand movement is visible, but not excessive. Deflation areas are small and localised, while dune breaches are shallow and episodic, without forming permanent channels. Sand supply remains sufficient to sustain dune development. Overall, this points to a balanced system in which aeolian processes are ongoing, but not leading to destabilisation.</p>
        <p>Ecological indicators (ECO-1 to ECO-5 = 4) confirm a very well-preserved vegetation structure. Vegetation cover exceeds 60%, with clear zonation and strong representation of both pioneer species (e.g. <tp:taxon-name><tp:taxon-name-part taxon-name-part-type="genus" reg="Ammophila">Ammophila</tp:taxon-name-part></tp:taxon-name>, Elytrigia) and typical psammophytes. Habitat diversity is high, including several dune habitat types, some of them of conservation priority. At the same time, the high score for habitat vulnerability reflects the presence of sensitive ecosystems that require continued protection.</p>
        <p>Pressure and risk indicators show more variable conditions. Marine exposure is high (PRS-1 = 1), indicating strong wave energy impact, minimal topographic protection and frequent storm overwash affecting the dune front. Despite this exposure, the system remains overall stable, suggesting a certain degree of resilience to marine forcing. In contrast, degradation risk (PRS-2 = 3) reflects moderate, but manageable pressures, such as localised trampling or minor erosion. Macro-litter pollution (PRS-3 = 3) is noticeable, but not dominant, while the presence of invasive alien species (PRS-4 = 2) indicates limited, but detectable ecological disturbance.</p>
        <p>Human footprint indicators reveal a mixed pattern. Permanent infrastructure and access paths (HFP-1 and HFP-2 = 4) are minimal or well managed, resulting in low direct physical disturbance. However, the recreational use intensity (HFP-4 = 2) indicates moderate visitor pressure, including localised trampling, particularly during peak seasons. Beach cleaning practices (HFP-3 = 3) suggest periodic mechanical intervention, which may have localised ecological effects.</p>
        <p>Legal and management indicators are generally favourable. Land ownership and access regulation (LEG-1 = 4) indicate a relatively well-controlled regime, while protected area status (LEG-2 = 3) reflects partial, but not fully effective conservation coverage. Cadastral recognition (LEG-3 = 4) and scenic value (LEG-4 = 4) confirm both formal recognition and high landscape quality. However, the absence of designation under EU or national restoration frameworks (LEG-5 = 0) stands out as a clear gap, indicating that the site is not yet integrated into strategic restoration planning.</p>
        <p>Scientific and conservation indicators (SCI-1, SCI-2, CON-1, CON-2 = 4) highlight the high research value and strategic importance of the system. The site includes well-preserved and representative geomorphological features, supports biodiversity of scientific interest and is suitable for long-term monitoring and conservation initiatives. The high feasibility of restoration further suggests that management interventions, if needed, could be effectively implemented.</p>
        <p>Overall, the Primorsko dune system can be considered a highly functional and largely intact coastal environment, where natural processes and ecological structure are well preserved. At the same time, the presence of moderate human pressures and gaps in policy integration highlights the need for targeted and proactive management. This case study demonstrates the ability of the EGDVI to capture both the high conservation value of dune systems and the more subtle pressures that may influence their long-term stability.</p>
      </sec>
    </sec>
    <sec sec-type="Discussion">
      <title>Discussion</title>
      <sec sec-type="Regional adaptation">
        <title>Regional adaptation</title>
        <p>Most existing dune assessment and valuation frameworks have been developed for macro- and mesotidal coastal environments, where tidal range exerts a dominant control on sediment exchange, dune morphology and disturbance regimes (<xref ref-type="bibr" rid="B13882327">Carter et al. 1991</xref>, <xref ref-type="bibr" rid="B13882929">Nordstrom 2000</xref>, <xref ref-type="bibr" rid="B13883068">Pye and Blott 2008</xref>). In such settings, frequent overwash, strong tidal currents and extensive intertidal zones fundamentally shape dune dynamics, often resulting in rapid morphological change and highly mobile vegetation patterns.</p>
        <p>In contrast, the Black Sea coast represents a microtidal, fetch-limited basin, where aeolian transport, wind-generated waves and episodic storm surges dominate coastal dynamics, while tidal forcing is negligible (<xref ref-type="bibr" rid="B13882746">Lin-Ye et al. 2018</xref>, <xref ref-type="bibr" rid="B13883147">Saprykina and Kuznetsov 2018</xref>). Under these conditions, dune systems tend to persist over longer timescales, sediment–vegetation feedbacks are more stable and successional stages often reach greater maturity than in high-energy tidal environments. These characteristics substantially influence both the expression of disturbance and the pathways of degradation and recovery, making direct transfer of assessment frameworks developed for mesotidal coasts problematic.</p>
        <p>The need for region-specific adaptation of dune assessment approaches has been explicitly recognised for other low-energy or semi-enclosed seas, including the Baltic and parts of the Mediterranean (<xref ref-type="bibr" rid="B13882270">Bagdanavičiūtė et al. 2015</xref>, <xref ref-type="bibr" rid="B13882388">Ciccarelli et al. 2017</xref>). The EGDVI directly addresses this need by integrating eco-geomorphological indicators tailored to the Black Sea context, together with dimensions reflecting anthropogenic pressure, legal protection, conservation feasibility and scientific value. These latter aspects are often treated implicitly or qualitatively in earlier indices (<xref ref-type="bibr" rid="B13882482">Doody 2013</xref>, <xref ref-type="bibr" rid="B13882223">Alessandrini et al. 2024</xref>). Their explicit quantification enables the index to capture not only the physical and ecological condition of dune systems, but also their institutional recognition and relevance for conservation planning.</p>
        <p>An additional rationale for focusing explicitly on dune systems rather than the full beach–dune continuum arises from contemporary coastal management practices in Bulgaria. Large sections of the active beach are subject to mechanical grooming, levelling and cleaning associated with concession management. Such practices suppress natural geomorphological signals and significantly alter ecological patterns, particularly those related to early successional vegetation (<xref ref-type="bibr" rid="B13882318">Carboni et al. 2009</xref>, <xref ref-type="bibr" rid="B13882777">Malavasi et al. 2014</xref>). Under these conditions, dune systems constitute a more stable, interpretable and ecologically meaningful unit for conservation-orientated assessment than beaches.</p>
        <p>This distinction is particularly important given that many existing indices explicitly assess the beach–dune system as a coupled unit, thereby integrating processes and pressures that are fundamentally altered by intensive beach management. By isolating dune habitats as the primary unit of analysis, the EGDVI avoids this methodological bias and ensures that assessed patterns reflect intrinsic eco-geomorphological processes rather than artefacts of beach maintenance.</p>
      </sec>
      <sec sec-type="Class-based interpretation of EGDVI results">
        <title>Class-based interpretation of EGDVI results</title>
        <p>The class structure derived from the EGDVI highlights that differences amongst dune systems along the Strandzha coast are not merely numerical, but reflect qualitatively distinct eco-geomorphological states. Rather than forming a continuous gradient, the systems cluster into value domains that correspond to recognisable combinations of physical integrity, ecological maturity and governance context. This pattern is consistent with the asymmetric score distribution and class-level statistics, which indicate both strong differentiation and pronounced regional heterogeneity.</p>
        <p>The use of fixed class thresholds facilitates interpretation beyond relative ranking, enabling the identification of meaningful transitions between conservation states. As such, the EGDVI classes provide a structured framework for linking observed eco-geomorphological conditions with management priorities and feasible intervention pathways.</p>
        <sec sec-type="Patterns and internal differentiation within classes">
          <title>Patterns and internal differentiation within classes</title>
          <p>Systems classified as Class A – Very High Eco-Geomorphological Value form a compact and coherent group. Their high mean scores and relatively limited internal dispersion indicate near-reference conditions across multiple dimensions. These systems consistently combine well-preserved geomorphological structure, advanced vegetation zonation, low anthropogenic pressure and favourable legal and management conditions. Importantly, no single indicator group dominates the final scores. Instead, very high values emerge from the cumulative alignment of geomorphological, ecological and governance-related factors.</p>
          <p>At the same time, internal differentiation within Class A indicates that near-reference conditions can be achieved through different, yet equally stable, configurations. Some systems attain very high values primarily through exceptional geomorphological development, while others are reinforced by strong legal protection, long-standing scientific recognition or high feasibility of conservation management. This diversity suggests that the very high eco-geomorphological value is not tied to a single idealised system type, but can arise through multiple pathways.</p>
          <p>The sole Class B – High Value system occupies a transitional position that is particularly informative from a management perspective. Its physical structure and ecological integrity closely resemble those of Class A systems, yet relatively modest pressures related to access and seasonal recreation constrain its overall performance. This distinction suggests that the boundary between high and very high value is often governed less by intrinsic physical limitations than by the effectiveness of governance and visitor management.</p>
          <p>Class C – Moderate Value represents the most extensive and regionally dominant group. Although all systems within this class retain functional dune elements and active eco-geomorphological processes, they differ substantially in their dominant constraints. In some cases, sustained human pressure is the primary limiting factor, while in others, reduced width, fragmentation or altered sediment connectivity play a more important role. Several systems occupy threshold positions close to the Class B boundary, indicating sensitivity to relatively small changes in pressure or management effectiveness. Despite a relatively narrow dispersion of scores, Class C should, therefore, be interpreted as a transitional domain rather than a uniform condition, encompassing multiple degradation and recovery trajectories.</p>
          <p>Class D – Low-Value systems exhibit reduced geomorphological functionality under persistent pressure, resulting in limited natural recovery potential. While not fully degraded, these systems lack the resilience required for spontaneous improvement and, therefore, require structured, intervention-orientated management.</p>
          <p>Finally, Class E – Degraded systems display uniformly low scores and minimal internal variability, reflecting an advanced and relatively homogeneous state of degradation. Near-complete loss of frontal dune structure, disrupted sediment pathways and sparse psammophytic vegetation indicate that degradation has progressed beyond reversible thresholds under current conditions.</p>
        </sec>
      </sec>
      <sec sec-type="Linking EGDVI classes to management and restoration frameworks">
        <title>Linking EGDVI classes to management and restoration frameworks</title>
        <p>The class structure produced by the EGDVI aligns closely with the logic underpinning contemporary management- and restoration-orientated dune frameworks. Unlike risk- or vulnerability-focused indices, which primarily prioritise sites based on exposure and sensitivity, the EGDVI reflects cumulative eco-geomorphological value and governance context, allowing management decisions to be guided not only by threat levels, but also by long-term conservation significance.</p>
        <p>Class A systems correspond to conservation-priority contexts where strict protection and long-term monitoring should take precedence over active intervention, consistent with best-practice guidance for intact dune systems (<xref ref-type="bibr" rid="B13882474">Denning et al. 2024</xref>). These systems align with the “protection-only” category of the ReDune framework (<xref ref-type="bibr" rid="B13882757">Lithgow et al. 2015</xref>) and with low-urgency categories of the Dune Restoration Potential (DRP) index (<xref ref-type="bibr" rid="B13882994">Pinna et al. 2022</xref>).</p>
        <p>Class B systems represent a preventative management tier in which relatively modest measures, such as access regulation, boardwalk installation and fencing, can prevent further degradation and facilitate upward transition. Evidence from comparable coastal settings indicates that such measures are effective in reducing trampling and informal path proliferation (<xref ref-type="bibr" rid="B13882938">Nordstrom 2015</xref>, <xref ref-type="bibr" rid="B13882474">Denning et al. 2024</xref>).</p>
        <p>Class C systems align most closely with restoration-orientated categories, in which targeted interventions can substantially improve eco-geomorphological conditions. Measures such as access control, invasive species management, cadastral clarification and re-vegetation with dune-building species are widely recognised as effective for moderately degraded systems (<xref ref-type="bibr" rid="B13882474">Denning et al. 2024</xref>). The numerical dominance of this class along the Strandzha coast highlights both the vulnerability of the regional dune landscape and its potential for meaningful improvement through coordinated management action.</p>
        <p>Class D and Class E systems correspond to progressively more demanding restoration and rehabilitation contexts. In these cases, recovery depends not only on ecological measures, but also on policy decisions, long-term institutional commitment and, in some instances, acceptance that full restoration may not be feasible (<xref ref-type="bibr" rid="B13882757">Lithgow et al. 2015</xref>, <xref ref-type="bibr" rid="B13882994">Pinna et al. 2022</xref>).</p>
      </sec>
      <sec sec-type="Comparative positioning of the EGDVI amongst dune assessment indices">
        <title>Comparative positioning of the EGDVI amongst dune assessment indices</title>
        <p>Over recent decades, dune assessment indices have evolved from checklist-based vulnerability approaches towards more integrated eco-geomorphological and management-orientated frameworks, although the majority remain primarily focused on condition, vulnerability or risk rather than on integrated value assessment (<xref ref-type="bibr" rid="B13886304">Bodéré et al. 1991</xref>, <xref ref-type="bibr" rid="B13886318">García-Mora et al. 2001</xref> etc.) - Table <xref ref-type="table" rid="T13883676">5</xref>. Eco-geomorphological indices such as the Coastal Dune Vulnerability Index (CDVI) represented an important methodological advance by explicitly integrating physical, ecological and anthropogenic dimensions (<xref ref-type="bibr" rid="B13882388">Ciccarelli et al. 2017</xref>, <xref ref-type="bibr" rid="B13882287">Bertoni et al. 2019</xref> etc.) - Table <xref ref-type="table" rid="T13883676">5</xref>. However, these frameworks rarely incorporate legal recognition, governance effectiveness or scientific significance as explicit and independently weighted quantitative components.</p>
        <p>The EGDVI advances the state of the art by explicitly integrating governance- and knowledge-related indicators alongside eco-geomorphological variables, thereby strengthening the science–policy interface (<xref ref-type="bibr" rid="B13882482">Doody 2013</xref>, <xref ref-type="bibr" rid="B13882223">Alessandrini et al. 2024</xref>). Unlike highly data-intensive multidimensional CVI or risk-based frameworks, which often require extensive ecological, socio-economic and spatial datasets and are best suited to large-scale applications (<xref ref-type="bibr" rid="B13882541">Furlan et al. 2021</xref>), the EGDVI is intentionally designed as a medium–high complexity, but standardised workflow, balancing analytical depth with operational applicability at the regional scale.</p>
        <p>Although the conservation status of dune habitats in Bulgaria is formally assessed under the reporting framework of the EU Habitats Directive (<xref ref-type="bibr" rid="B13882409">Council Directive 92/43/EEC 1992</xref>), these assessments are often associated with considerable uncertainty due to broader challenges in Natura 2000 implementation, including limited standardised field-based monitoring and methodological inconsistencies (<xref ref-type="bibr" rid="B14211912">Tzonev et al. 2023</xref>). Consequently, national-scale evaluations may not fully reflect the spatial heterogeneity and site-specific ecological conditions of coastal dune systems.</p>
        <p>In this context, the Eco-Geomorphological Dune Value Index (EGDVI) is not directly equivalent to the conservation status categories applied under Article 17 (<xref ref-type="bibr" rid="B13882409">Council Directive 92/43/EEC 1992</xref>) reporting, as it follows a different conceptual and methodological approach. Rather than focusing solely on habitat conservation status, the EGDVI provides an integrative site-scale assessment that combines geomorphological integrity, vegetation structure, anthropogenic pressures, legal and management characteristics, scientific significance and restoration potential within a unified framework.</p>
        <p>As such, the EGDVI should be viewed as complementary to Article 17 assessments (<xref ref-type="bibr" rid="B13882409">Council Directive 92/43/EEC 1992</xref>), offering an additional evidence-based perspective that may help reduce uncertainty and support more spatially explicit and operational conservation evaluations in future reporting cycles.</p>
        <p>By combining fixed classification thresholds with sensitivity analysis and by explicitly linking classes to management and restoration pathways, the EGDVI moves beyond descriptive assessment towards actionable conservation planning. In this sense, it synthesises the strengths of previous indices while introducing a governance-aware, value-based and class-orientated framework tailored to low-energy, microtidal coastal environments such as the Black Sea.</p>
      </sec>
    </sec>
    <sec sec-type="Conclusions">
      <title>Conclusions</title>
      <p>The Eco-Geomorphological Dune Value Index (EGDVI) provides an integrated, standardised and repeatable framework for assessing the conservation value of coastal dune systems by combining geomorphological structure, ecological condition, anthropogenic pressure, legal protection and scientific value within a single operational tool. Its application to 16 dune systems along the Strandzha sector of the Bulgarian Black Sea coast produced total scores ranging from 154.0 to 364.5 (out of a theoretical maximum of 400) and resulted in a clear five-class structure under fixed classification thresholds.</p>
      <p>Beyond individual site classification, the results allow a synthetic regional interpretation of dune condition along the Strandzha coast. The mean EGDVI score for all assessed systems is 266.1, with a median of 261.2, placing the overall regional condition within the moderate-value class (Class C). This regional average does not represent a new index, but rather a descriptive benchmark that captures the dominant eco-geomorphological state of the coastline as a whole.</p>
      <p>The regional mean highlights that, despite the presence of several near-reference dune systems with very high eco-geomorphological value, the Strandzha coast is predominantly characterised by transitional dune systems. These systems retain functional geomorphological and ecological components, but are subject to persistent pressures and structural constraints that limit their overall performance. As a result, a small number of highly preserved sites does not offset the broader regional pattern of moderate condition.</p>
      <p>The class-based structure of the EGDVI further demonstrates that similar overall values may arise from different combinations of drivers and constraints. Very high and high scores reflect not only geomorphological integrity, but also the cumulative influence of governance-related factors such as legal protection, conservation feasibility and scientific recognition. Conversely, moderate and low values may be driven by sustained human pressure, reduced sediment connectivity, or spatial fragmentation. This diagnostic capacity distinguishes the EGDVI from simple ranking approaches and enhances its relevance for management-orientated interpretation.</p>
      <p>By explicitly incorporating indicators adapted to the microtidal, fetch-limited dynamics of the Black Sea, the EGDVI addresses a key limitation of many existing dune assessment and vulnerability indices developed primarily for macro- or mesotidal coasts. The framework’s modular structure, transparent weighting scheme and fixed class thresholds support repeatability in long-term monitoring and conceptual transferability to other low-energy or semi-enclosed coastal environments, provided that indicator thresholds are locally calibrated.</p>
      <p>Beyond site comparison, the EGDVI functions as a diagnostic and decision-support tool, identifying priority areas for conservation, highlighting systems where preventative management may avert further decline and distinguishing sites where restoration or rehabilitation measures are likely to be most effective. The explicit inclusion of legal protection and scientific value strengthens the science–policy interface by ensuring that institutional recognition and research relevance are adequately reflected in conservation planning.</p>
      <p>Current limitations relate primarily to data availability and to the need for calibration beyond the Black Sea region. Nevertheless, the EGDVI provides a robust baseline for adaptive management and long-term monitoring of dune systems. Future refinements through structured expert elicitation, the integration of time-series observations and the use of emerging mapping technologies, such as UAS- and LiDAR-based analyses, including machine-learning approaches, are expected to further enhance its analytical resolution and operational applicability.</p>
      <p>In summary, the EGDVI represents a substantive methodological advance in coastal dune assessment by integrating eco-geomorphological conditions, anthropogenic pressures and governance dimensions into a single, coherent framework. Its application along the Strandzha coast demonstrates both its diagnostic capacity at the site level and its ability to provide a meaningful regional synthesis, offering a transferable model for the assessment and management of dune systems in other microtidal coastal environments.</p>
    </sec>
  </body>
  <back>
    <ack>
      <title>Acknowledgements</title>
      <p>The index was developed under the project “<bold><italic>Mapping and Spatiotemporal Analysis of Beach-Dune Systems on the Southern Bulgarian Black Sea Coast: Evolution, Anthropogenic Pressure and Ecological Risks to Dune Habitats (MapBGBeachDune)</italic></bold>”. Grant № КП-06-H84/5-16.12.2024 under Competition for financial support of basic research projects of the Bulgarian National Science Fund.</p>
      <p>The authors would also like to express sincere gratitude to Dr. Petko Bozhkov, Dr. Ilia Tamburadzhiev, Todor Lambev, Dobroslav Dechev, Nadezhda Dimitrova, Lyubomir Rasovski, Nina Tzvetkova and Chavdar Gussev for their active participation in field campaigns and expedition-based investigations, as well as to the students Nadezhda Dimitrova, Nina Tsvetkova and Lyubomir Rasovski for their valuable assistance during fieldwork and data collection.</p>
    </ack>
    <sec sec-type="Author contributions">
      <title>Author contributions</title>
      <p><italic>Bogdan K. Prodanov</italic>: Conceptualisation; Methodology; Index development; Funding acquisition; Project administration; Resources; Software; Field investigation; Geomorphological and ecological data collection; Data curation; Formal analysis; Validation; Visualisation; Writing – original draft; Writing – review and editing; <italic>Radoslava I. Bekova</italic>: Software; Methodology; Field investigation; Geomorphological and ecological data collection; Data curation; Formal analysis; Validation; Visualisation; Writing – original draft; Writing – review and editing; <italic>Magdalena I. Valcheva</italic>: Methodology; Field investigation; Ecological data collection; Data curation; Formal analysis; Validation; Writing – original draft; Writing – review and editing; <italic>Rossen Tzonev</italic>: Field investigation; Writing – review and editing; Critical note; <italic>Ahinora G. Baltakova</italic>: Field investigation; Geomorphological data collection; Data curation; Formal analysis; Validation; Writing – review and editing.</p>
    </sec>
    <sec sec-type="Conflicts of interest">
      <title>Conflicts of interest</title>
      <p>No conflict of interest to declare</p>
      <p>Disclaimer: This article is (co-)authored by any of the Editors-in-Chief, Managing Editors or their deputies in this journal.</p>
    </sec>
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  </back>
  <floats-group>
    <fig id="F13883313" position="float" orientation="portrait">
      <object-id content-type="arpha">45E27FE1-8A47-5BF7-884E-C02EEEA58B0C</object-id>
      
      <label>Figure 1.</label>
      <caption>
        <p>Location and type of coastal dune system along Strandzha Black Sea Coast, Bulgaria.</p>
      </caption>
      <graphic xlink:href="oneeco-11-e189660-g001.png" position="float" id="oo_1547668.png" orientation="portrait" xlink:type="simple">
        <uri content-type="original_file">https://binary.pensoft.net/fig/1547668</uri>
      </graphic>
    </fig>
    <fig id="F13883315" position="float" orientation="portrait">
      <object-id content-type="arpha">C451DF5A-A19B-5CB8-A47C-44D1719A3455</object-id>
      
      <label>Figure 2.</label>
      <caption>
        <p>Aerophotographs of beach-dune systems (1-6) along Strandzha coast, Bulgaria (author: Bogdan Prodanov), <italic>see location on</italic> <bold><italic>Fig. 1.</italic></bold></p>
      </caption>
      <graphic xlink:href="oneeco-11-e189660-g002.png" position="float" id="oo_1535062.png" orientation="portrait" xlink:type="simple">
        <uri content-type="original_file">https://binary.pensoft.net/fig/1535062</uri>
      </graphic>
    </fig>
    <fig id="F13883317" position="float" orientation="portrait">
      <object-id content-type="arpha">9859EA32-16BB-545F-A1C7-6E0B8815BD2E</object-id>
      
      <label>Figure 3.</label>
      <caption>
        <p>Aerophotographs of beach-dune systems (7, 10-13,16) along Strandzha coast, Bulgaria (author: Bogdan Prodanov), <italic>see location on <bold>Fig. 1</bold></italic>.</p>
      </caption>
      <graphic xlink:href="oneeco-11-e189660-g003.png" position="float" id="oo_1535063.png" orientation="portrait" xlink:type="simple">
        <uri content-type="original_file">https://binary.pensoft.net/fig/1535063</uri>
      </graphic>
    </fig>
    <fig id="F13883319" position="float" orientation="portrait">
      <object-id content-type="arpha">0F20228F-2B47-53F8-A048-88E71CD3BD44</object-id>
      
      <label>Figure 4.</label>
      <caption>
        <p>Spatial distribution, total scores and classification of coastal dune systems along the Strandzha Black Sea Coast (Bulgaria) according to the EGDVI.</p>
      </caption>
      <graphic xlink:href="oneeco-11-e189660-g004.png" position="float" id="oo_1535064.png" orientation="portrait" xlink:type="simple">
        <uri content-type="original_file">https://binary.pensoft.net/fig/1535064</uri>
      </graphic>
    </fig>
    <fig id="F14194734" position="float" orientation="portrait">
      <object-id content-type="arpha">1520B6E3-D46C-57E7-9A58-5457F7E5A40C</object-id>
      
      <label>Figure 5.</label>
      <caption>
        <p>Aerophotography of Primorsko (Stomopolu-Perla) beach-dune system.</p>
      </caption>
      <graphic xlink:href="oneeco-11-e189660-g005.jpg" position="float" id="oo_1621042.JPG" orientation="portrait" xlink:type="simple">
        <uri content-type="original_file">https://binary.pensoft.net/fig/1621042</uri>
      </graphic>
    </fig>
    <table-wrap id="T13883321" position="float" orientation="portrait">
      <label>Table 1.</label>
      <caption>
        <p>Categories, indicators and measurement methods for evaluating the EGDVI (<italic>A full version of indicators, definitions and scoring criteria is provided in Suppl. materials <xref ref-type="supplementary-material" rid="S13883629">1</xref>, <xref ref-type="supplementary-material" rid="S13886327">2</xref></italic>).</p>
      </caption>
      <table rules="all" border="1">
        <tbody>
          <tr>
            <td rowspan="1" colspan="1">Category</td>
            <td rowspan="1" colspan="1">Indicator Abr.</td>
            <td rowspan="1" colspan="1">Indicator(<bold><italic>scores: 0-4 in Appendix 1</italic></bold>)</td>
            <td rowspan="1" colspan="1">Weight (%)</td>
            <td rowspan="1" colspan="1">Max. scores</td>
            <td rowspan="1" colspan="1">Relevant source/Available scientific data<break/>(<italic>For pilot case study</italic>)</td>
            <td rowspan="1" colspan="1">Methods, Measurement, and Survey</td>
          </tr>
        </tbody>
        <tbody>
          <tr>
            <td rowspan="5" colspan="1">Geomorphology</td>
            <td rowspan="1" colspan="1">
              <bold>GM-1</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Geomorphological Diversity</italic>
            </td>
            <td rowspan="1" colspan="1">
              <italic>6</italic>
            </td>
            <td rowspan="1" colspan="1">24</td>
            <td rowspan="5" colspan="1">(<italic><xref ref-type="bibr" rid="B13883036">Prodanov et al. 2023</xref>, <xref ref-type="bibr" rid="B13883046">Prodanov et al. 2025</xref>, <xref ref-type="bibr" rid="B13883012">Prodanov et al. 2026</xref>)</italic></td>
            <td rowspan="5" colspan="1">
              <italic>Field work/ GIS/UAS-Photogrammetry, LiDAR techniques</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>GM-2</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Dune Height (m)</italic>
            </td>
            <td rowspan="1" colspan="1">6</td>
            <td rowspan="1" colspan="1">24</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>GM-3</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Dune System Width (m)</italic>
            </td>
            <td rowspan="1" colspan="1">6</td>
            <td rowspan="1" colspan="1">24</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>GM-4</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Hydrogeomorphic Connectivity (Dune–Wetland Interface)</italic>
            </td>
            <td rowspan="1" colspan="1">6</td>
            <td rowspan="1" colspan="1">24</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>GM-5</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Connectivity/Fragmentation of the Dune System</italic>
            </td>
            <td rowspan="1" colspan="1">6</td>
            <td rowspan="1" colspan="1">24</td>
          </tr>
          <tr>
            <td rowspan="4" colspan="1">Aeolian Dynamics</td>
            <td rowspan="1" colspan="1">
              <bold>AD-1</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Extent of blowouts (% of the dune system)</italic>
            </td>
            <td rowspan="1" colspan="1">1.5</td>
            <td rowspan="1" colspan="1">6</td>
            <td rowspan="4" colspan="1">(<italic><xref ref-type="bibr" rid="B13883036">Prodanov et al. 2023</xref>, <xref ref-type="bibr" rid="B13883012">Prodanov et al. 2026</xref>)</italic></td>
            <td rowspan="4" colspan="1">
              <italic>Field work/ GIS</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>AD-2</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Deflation and sand mobility</italic>
            </td>
            <td rowspan="1" colspan="1">1.5</td>
            <td rowspan="1" colspan="1">6</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>AD-3</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Dune breaches – depth as % of dune height</italic>
            </td>
            <td rowspan="1" colspan="1">1.5</td>
            <td rowspan="1" colspan="1">6</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>AD-4</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Sand supply input from wind</italic>
            </td>
            <td rowspan="1" colspan="1">1.5</td>
            <td rowspan="1" colspan="1">6</td>
          </tr>
          <tr>
            <td rowspan="5" colspan="1">Ecology</td>
            <td rowspan="1" colspan="1">
              <bold>ECO-1</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Vegetation Integrity</italic>
            </td>
            <td rowspan="1" colspan="1">5.5</td>
            <td rowspan="1" colspan="1">26</td>
            <td rowspan="5" colspan="1">(<italic><xref ref-type="bibr" rid="B13883220">Tzonev et al. 2005</xref>, <xref ref-type="bibr" rid="B13882609">Gussev and Tzonev 2014</xref>, <xref ref-type="bibr" rid="B13883237">Valcheva et al. 2020</xref>, <xref ref-type="bibr" rid="B13883255">Valcheva et al. 2021</xref>)</italic><break/><italic>Cadastral and specialised maps</italic>: <ext-link ext-link-type="uri" xlink:href="https://kais.cadastre.bg/en/Map">https://kais.cadastre.bg/en/Map</ext-link><break/>Suppl. material <xref ref-type="supplementary-material" rid="S13886328">3</xref></td>
            <td rowspan="5" colspan="1">
              <italic>Field work/ GIS/Laboratory</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>ECO-2</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Presence of Pioneer Dune Species</italic>
            </td>
            <td rowspan="1" colspan="1">5.5</td>
            <td rowspan="1" colspan="1">26</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>ECO-3</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Presence of Typical Psammophytes</italic>
            </td>
            <td rowspan="1" colspan="1">5</td>
            <td rowspan="1" colspan="1"/>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>ECO-4</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Habitat Diversity</italic>
            </td>
            <td rowspan="1" colspan="1">5</td>
            <td rowspan="1" colspan="1">26</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>ECO-5</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Habitat Vulnerability</italic>
            </td>
            <td rowspan="1" colspan="1">5</td>
            <td rowspan="1" colspan="1">26</td>
          </tr>
          <tr>
            <td rowspan="4" colspan="1">Pressure/Risk/Stress</td>
            <td rowspan="1" colspan="1">
              <bold>PRS-1</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Marine Exposure</italic>
            </td>
            <td rowspan="1" colspan="1">2</td>
            <td rowspan="1" colspan="1">10</td>
            <td rowspan="4" colspan="1">(<italic><xref ref-type="bibr" rid="B13883237">Valcheva et al. 2020</xref>, <xref ref-type="bibr" rid="B13883255">Valcheva et al. 2021</xref>, <xref ref-type="bibr" rid="B13889458">Bekova and Prodanov 2023</xref>, <xref ref-type="bibr" rid="B13889476">Bekova and Prodanov 2024</xref>, <xref ref-type="bibr" rid="B13889467">Prodanov and Bekova 2023</xref>, <xref ref-type="bibr" rid="B13889448">Valchev et al. 2023</xref>)</italic>;<break/><italic>Suppl. material <xref ref-type="supplementary-material" rid="S13886328">3</xref></italic></td>
            <td rowspan="4" colspan="1">
              <italic>Field work/ GIS/Laboratory</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>PRS-2</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Degradation Risk</italic>
            </td>
            <td rowspan="1" colspan="1">2.5</td>
            <td rowspan="1" colspan="1">10</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>PRS-3</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Macro-litter Pollution</italic>
            </td>
            <td rowspan="1" colspan="1">1.5</td>
            <td rowspan="1" colspan="1">10</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>PRS-4</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Presence of Invasive Alien Plants</italic>
            </td>
            <td rowspan="1" colspan="1">1.5</td>
            <td rowspan="1" colspan="1"/>
          </tr>
          <tr>
            <td rowspan="4" colspan="1">Human Footprint Pressure (HFP)</td>
            <td rowspan="1" colspan="1">
              <bold>HFP-1</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Permanent infrastructure on dunes (% of surface)</italic>
            </td>
            <td rowspan="1" colspan="1">2.5</td>
            <td rowspan="1" colspan="1">10</td>
            <td rowspan="4" colspan="1"/>
            <td rowspan="4" colspan="1">
              <italic>Field work/ GIS</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>HFP-2</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Access paths/trails (% of frontal dune)</italic>
            </td>
            <td rowspan="1" colspan="1">2.5</td>
            <td rowspan="1" colspan="1">10</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>HFP-3</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Beach cleaning frequency and width</italic>
            </td>
            <td rowspan="1" colspan="1">2.5</td>
            <td rowspan="1" colspan="1">10</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>HFP-4</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Recreational use intensity (visitor pressure/frequency, observed trampling)</italic>
            </td>
            <td rowspan="1" colspan="1">2</td>
            <td rowspan="1" colspan="1">8</td>
          </tr>
          <tr>
            <td rowspan="5" colspan="1">Legal/Protection Status/Management</td>
            <td rowspan="1" colspan="1">
              <bold>LEG-1</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Land Ownership &amp; Access Pressure</italic>
            </td>
            <td rowspan="1" colspan="1">2</td>
            <td rowspan="1" colspan="1">8</td>
            <td rowspan="5" colspan="1"><italic>Cadastral and specialised maps</italic>: <ext-link ext-link-type="uri" xlink:href="https://kais.cadastre.bg/en/Map">https://kais.cadastre.bg/en/Map</ext-link>; <break/>(<italic><xref ref-type="bibr" rid="B13882892">Mooser et al. 2022</xref>)</italic></td>
            <td rowspan="5" colspan="1">
              <italic>GIS</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>LEG-2</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Protected Area Status</italic>
            </td>
            <td rowspan="1" colspan="1">2</td>
            <td rowspan="1" colspan="1">8</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>LEG-3</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Cadastral Recognition</italic>
            </td>
            <td rowspan="1" colspan="1">2</td>
            <td rowspan="1" colspan="1">8</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>LEG-4</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Scenic Value</italic>
            </td>
            <td rowspan="1" colspan="1">2</td>
            <td rowspan="1" colspan="1">8</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>LEG-5</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Policy Designation under EU/National Restoration Frameworks</italic>
            </td>
            <td rowspan="1" colspan="1">2</td>
            <td rowspan="1" colspan="1">8</td>
          </tr>
          <tr>
            <td rowspan="2" colspan="1">Scientific importance</td>
            <td rowspan="1" colspan="1">
              <bold>SCI-1</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Geomorphological Scientific Value</italic>
            </td>
            <td rowspan="1" colspan="1">2</td>
            <td rowspan="1" colspan="1">8</td>
            <td rowspan="2" colspan="1">(<italic><xref ref-type="bibr" rid="B13883036">Prodanov et al. 2023</xref>, <xref ref-type="bibr" rid="B13883012">Prodanov et al. 2026</xref>, <xref ref-type="bibr" rid="B13883220">Tzonev et al. 2005</xref>, <xref ref-type="bibr" rid="B13882609">Gussev and Tzonev 2014</xref>, <xref ref-type="bibr" rid="B13883237">Valcheva et al. 2020</xref>, <xref ref-type="bibr" rid="B13883255">Valcheva et al. 2021</xref>)</italic>;<break/><italic>Suppl. material <xref ref-type="supplementary-material" rid="S13886328">3</xref></italic></td>
            <td rowspan="2" colspan="1">
              <italic>Field work/ GIS</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>SCI-2</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Ecological Scientific Value</italic>
            </td>
            <td rowspan="1" colspan="1">2</td>
            <td rowspan="1" colspan="1">8</td>
          </tr>
          <tr>
            <td rowspan="2" colspan="1">Nature Conservation</td>
            <td rowspan="1" colspan="1">
              <bold>CON-1</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Strategic Conservation Value</italic>
            </td>
            <td rowspan="1" colspan="1">4</td>
            <td rowspan="1" colspan="1">16</td>
            <td rowspan="2" colspan="1">(<xref ref-type="bibr" rid="B13883220">Tzonev et al. 2005</xref>, <xref ref-type="bibr" rid="B13882609">Gussev and Tzonev 2014</xref>, <xref ref-type="bibr" rid="B13882588">Gussev 2015</xref>, <xref ref-type="bibr" rid="B13883175">Tzonev 2015a</xref>, <xref ref-type="bibr" rid="B13883184">Tzonev 2015b</xref>, <xref ref-type="bibr" rid="B13883193">Tzonev 2015c</xref>, <xref ref-type="bibr" rid="B13883202">Tzonev 2015d</xref>, <xref ref-type="bibr" rid="B13883211">Tzonev 2015e</xref>, <xref ref-type="bibr" rid="B13883229">Tzonev and Gussev 2015</xref>, <xref ref-type="bibr" rid="B13883237">Valcheva et al. 2020</xref>, <xref ref-type="bibr" rid="B13883255">Valcheva et al. 2021</xref>)</td>
            <td rowspan="2" colspan="1">
              <italic>Field work/ GIS</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>CON-2</bold>
            </td>
            <td rowspan="1" colspan="1">
              <italic>Feasibility of Restoration Implementation</italic>
            </td>
            <td rowspan="1" colspan="1">3</td>
            <td rowspan="1" colspan="1">12</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="4"><bold><italic>Total weight = 100</italic></bold>% </td>
            <td rowspan="1" colspan="3">
              <bold>
                <italic>400 – Maximum total score</italic>
              </bold>
            </td>
          </tr>
        </tbody>
      </table>
    </table-wrap>
    <table-wrap id="T13883322" position="float" orientation="portrait">
      <label>Table 2.</label>
      <caption>
        <p>Final EGDVI classification scheme, score ranges and interpretation of conservation value for coastal dune systems.</p>
      </caption>
      <table rules="all" border="1">
        <tbody>
          <tr>
            <td rowspan="1" colspan="1">Class</td>
            <td rowspan="1" colspan="1">Range (scores)</td>
            <td rowspan="1" colspan="1">Definition of Eco-Geomorphological Value</td>
            <td rowspan="1" colspan="1">
              <bold>Interpretation</bold>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">
              <bold>A – Very High</bold>
            </td>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">≥ <bold>320</bold></td>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">
              <bold>Very High Value</bold>
            </td>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb; vertical-align:top">A pristine or near-pristine coastal dune system exhibiting exceptional ecological quality, structural complexity and geomorphological integrity. The site includes well-developed zonation, high vegetation integrity, active dune dynamics and full legal or scientific recognition. It holds strategic importance at a national or international level (e.g. part of <italic>Natura 2000, Protected area</italic>).</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#c6e0b4">
              <bold>B – High</bold>
            </td>
            <td rowspan="1" colspan="1" style="background-color:#c6e0b4">
              <bold>319-280</bold>
            </td>
            <td rowspan="1" colspan="1" style="background-color:#c6e0b4">
              <bold>High Value</bold>
            </td>
            <td rowspan="1" colspan="1" style="background-color:#c6e0b4; vertical-align:top">A well-preserved dune system with intact structure and ecological functions, though potentially exposed to minor or seasonal pressures (e.g. <italic>tourism, grazing or erosion</italic>). Zonation is mostly retained, with good vegetation cover and geomorphological features. The system is suitable for proactive conservation, low-impact use and landscape-scale planning.</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">
              <bold>C – Moderate</bold>
            </td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">
              <bold>279-220</bold>
            </td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">
              <bold>Moderate Value</bold>
            </td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00; vertical-align:top">A partially degraded or structurally fragmented dune system, typically affected by moderate anthropogenic pressures, infrastructure or reduced habitat diversity. While natural values persist, ecological functionality is declining. The site still offers viable potential for targeted conservation, restoration and improved management.</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#ffc000">
              <bold>D – Low</bold>
            </td>
            <td rowspan="1" colspan="1" style="background-color:#ffc000">
              <bold>219-160</bold>
            </td>
            <td rowspan="1" colspan="1" style="background-color:#ffc000">
              <bold>Low Value</bold>
            </td>
            <td rowspan="1" colspan="1" style="background-color:#ffc000; vertical-align:top">A highly impacted dune system with significant structural and functional degradation. Key ecological components are disrupted or absent. The system may be isolated, artificially stabilised or affected by uncontrolled access, development or pollution.</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#ff7c80">
              <bold>E – Degraded</bold>
            </td>
            <td rowspan="1" colspan="1" style="background-color:#ff7c80">≤ <bold>159</bold></td>
            <td rowspan="1" colspan="1" style="background-color:#ff7c80">
              <bold>Degraded/Critical</bold>
            </td>
            <td rowspan="1" colspan="1" style="background-color:#ff7c80; vertical-align:top">An ecologically and geomorphologically collapsed dune system, often entirely disconnected from natural dynamics. Characterised by artificial surfaces, severe pollution, total loss of vegetation structure and unregulated development. Represents an ecological emergency with negligible residual natural value unless urgent action is taken.</td>
          </tr>
        </tbody>
      </table>
    </table-wrap>
    <table-wrap id="T13883325" position="float" orientation="portrait">
      <label>Table 3.</label>
      <caption>
        <p>Evaluating EGDVI of dune systems.</p>
      </caption>
      <table rules="all" border="1">
        <tbody>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>Dune System</bold>
            </td>
            <td rowspan="1" colspan="1">
              <bold>Total</bold>
              <break/>
              <bold>score</bold>
            </td>
            <td rowspan="1" colspan="1">
              <bold>Classes</bold>
            </td>
          </tr>
        </tbody>
        <tbody>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">01 Primorsko (Stomopolu - Perla)</td>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">364.5</td>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">
              <italic>Class A – Very High Eco-Geomorphological Value</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">02 Primorsko (International Youth Centre)</td>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">357</td>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">
              <italic>Class A – Very High Eco-Geomorphological Value</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#a8d08d">03 Atliman</td>
            <td rowspan="1" colspan="1" style="background-color:#a8d08d">283.5</td>
            <td rowspan="1" colspan="1" style="background-color:#a8d08d">
              <italic>Class B – High Value</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">04 Kiten</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">264</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">
              <italic>Class C – Moderate Value</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">05 Koral and Camping Yug</td>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">345</td>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">
              <italic>Class A – Very High Eco-Geomorphological Value</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#ffc000">06 Lozenets</td>
            <td rowspan="1" colspan="1" style="background-color:#ffc000">201,5</td>
            <td rowspan="1" colspan="1" style="background-color:#ffc000">
              <italic>Class D – Low Value</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">07 Oazis</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">241</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">
              <italic>Class C – Moderate Value</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#ff7c80">08 Arapya</td>
            <td rowspan="1" colspan="1" style="background-color:#ff7c80">154</td>
            <td rowspan="1" colspan="1" style="background-color:#ff7c80">
              <italic>Class E – Degraded</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#ff7c80">09 Tsarevo (Popski plazh)</td>
            <td rowspan="1" colspan="1" style="background-color:#ff7c80">157.5</td>
            <td rowspan="1" colspan="1" style="background-color:#ff7c80">
              <italic>Class E – Degraded</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">10 Nestinarka</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">251.5</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">
              <italic>Class C – Moderate Value</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">11 Ahtopol</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">260.5</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">
              <italic>Class C – Moderate Value</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">12 Veleka River Mouth</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">278</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">
              <italic>Class C – Moderate Value</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">13 Butamyata</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">262</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">
              <italic>Class C – Moderate Value</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">14 Lipite</td>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">323</td>
            <td rowspan="1" colspan="1" style="background-color:#8eaadb">
              <italic>Class A – Very High Eco-Geomorphological Value</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">15 Listi</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">258</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">
              <italic>Class C – Moderate Value</italic>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">16 Silistar</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">256</td>
            <td rowspan="1" colspan="1" style="background-color:#ffff00">
              <italic>Class C – Moderate Value</italic>
            </td>
          </tr>
        </tbody>
      </table>
    </table-wrap>
    <table-wrap id="T13883326" position="float" orientation="portrait">
      <label>Table 4.</label>
      <caption>
        <p>Summary statistics of EGDVI scores by class.</p>
      </caption>
      <table rules="all" border="1">
        <tbody>
          <tr>
            <td rowspan="1" colspan="1">
              <bold>Class</bold>
            </td>
            <td rowspan="1" colspan="1">
              <bold>n</bold>
            </td>
            <td rowspan="1" colspan="1">
              <bold>Mean</bold>
            </td>
            <td rowspan="1" colspan="1">
              <bold>Std. Dev.</bold>
            </td>
            <td rowspan="1" colspan="1">
              <bold>Median</bold>
            </td>
            <td rowspan="1" colspan="1">
              <bold>Q1</bold>
            </td>
            <td rowspan="1" colspan="1">
              <bold>Q3</bold>
            </td>
            <td rowspan="1" colspan="1">
              <bold>IQR</bold>
            </td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">A – Very High</td>
            <td rowspan="1" colspan="1">4.00</td>
            <td rowspan="1" colspan="1">347.40</td>
            <td rowspan="1" colspan="1">15.70</td>
            <td rowspan="1" colspan="1">351.00</td>
            <td rowspan="1" colspan="1">339.50</td>
            <td rowspan="1" colspan="1">358.90</td>
            <td rowspan="1" colspan="1">19.40</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">B – High</td>
            <td rowspan="1" colspan="1">1.00</td>
            <td rowspan="1" colspan="1">283.50</td>
            <td rowspan="1" colspan="1">–</td>
            <td rowspan="1" colspan="1">283.50</td>
            <td rowspan="1" colspan="1">–</td>
            <td rowspan="1" colspan="1">–</td>
            <td rowspan="1" colspan="1">–</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">C – Moderate</td>
            <td rowspan="1" colspan="1">8.00</td>
            <td rowspan="1" colspan="1">258.90</td>
            <td rowspan="1" colspan="1">9.90</td>
            <td rowspan="1" colspan="1">259.20</td>
            <td rowspan="1" colspan="1">254.90</td>
            <td rowspan="1" colspan="1">262.50</td>
            <td rowspan="1" colspan="1">7.60</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">D – Low</td>
            <td rowspan="1" colspan="1">1.00</td>
            <td rowspan="1" colspan="1">201.50</td>
            <td rowspan="1" colspan="1">–</td>
            <td rowspan="1" colspan="1">201.50</td>
            <td rowspan="1" colspan="1">–</td>
            <td rowspan="1" colspan="1">–</td>
            <td rowspan="1" colspan="1">–</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">E – Degraded</td>
            <td rowspan="1" colspan="1">2.00</td>
            <td rowspan="1" colspan="1">155.80</td>
            <td rowspan="1" colspan="1">1.80</td>
            <td rowspan="1" colspan="1">155.80</td>
            <td rowspan="1" colspan="1">154.90</td>
            <td rowspan="1" colspan="1">156.60</td>
            <td rowspan="1" colspan="1">1.80</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">All systems<break/>(Strandzha coast)</td>
            <td rowspan="1" colspan="1">16.00</td>
            <td rowspan="1" colspan="1">266.10</td>
            <td rowspan="1" colspan="1">59.90</td>
            <td rowspan="1" colspan="1">261.20</td>
            <td rowspan="1" colspan="1">248.90</td>
            <td rowspan="1" colspan="1">293.40</td>
            <td rowspan="1" colspan="1">44.50</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="8">*Quartiles and IQR are not reported for classes with n = 1, as these statistics are<break/>not meaningful for single observations.</td>
          </tr>
        </tbody>
      </table>
    </table-wrap>
    <table-wrap id="T13883676" position="float" orientation="portrait">
      <label>Table 5.</label>
      <caption>
        <p>Conceptual and methodological frameworks for coastal dune assessment.</p>
      </caption>
      <table rules="all" border="0">
        <tbody>
          <tr>
            <td rowspan="1" colspan="1">Index / Framework</td>
            <td rowspan="1" colspan="1">Focus and indicators</td>
            <td rowspan="1" colspan="1">Data requirements /<break/>complexity</td>
            <td rowspan="1" colspan="1">Regional applicability</td>
            <td rowspan="1" colspan="1">Key methodological advance relative to EGDVI</td>
            <td rowspan="1" colspan="1">Reference</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">Checklist-based dune vulnerability assessment (Actual/Real Vulnerability Index)</td>
            <td rowspan="1" colspan="1">Structured checklist of geomorphological condition, surface stability, vegetation cover, human use and management pressure; summed scores yield an Actual/Real Vulnerability Index</td>
            <td rowspan="1" colspan="1">Medium–High (field observations + expert scoring via checklist)</td>
            <td rowspan="1" colspan="1">Atlantic coasts (France, Wales)</td>
            <td rowspan="1" colspan="1">EGDVI shifts from single-dimension vulnerability scoring to multi-domain value assessment, including legal and scientific dimensions</td>
            <td rowspan="1" colspan="1">(<xref ref-type="bibr" rid="B13886304">Bodéré et al. 1991</xref>)</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">Dune Vulnerability Index (DVI)</td>
            <td rowspan="1" colspan="1">Geomorphology, vegetation cover, marine influence, aeolian dynamics and human impact; five partial indices combined into a global vulnerability index</td>
            <td rowspan="1" colspan="1">Medium–High (field surveys, GIS, expert ranking)</td>
            <td rowspan="1" colspan="1">SW Iberian Peninsula</td>
            <td rowspan="1" colspan="1">EGDVI replaces vulnerability-only logic with value-based assessment and adds governance-related indicators</td>
            <td rowspan="1" colspan="1">(<xref ref-type="bibr" rid="B13886318">García-Mora et al. 2001</xref>)</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">H′dune / E-dune vegetation diversity indices</td>
            <td rowspan="1" colspan="1">Vegetation diversity, evenness and disturbance sensitivity as proxies for dune condition</td>
            <td rowspan="1" colspan="1">Low–Medium (vegetation plots)</td>
            <td rowspan="1" colspan="1">Northern and Central Europe</td>
            <td rowspan="1" colspan="1">EGDVI complements biodiversity-based indicators with geomorphological structure and anthropogenic pressure</td>
            <td rowspan="1" colspan="1">(<xref ref-type="bibr" rid="B13882579">Grunewald and Schubert 2007</xref>)</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">Multiscale dune conservation status assessment</td>
            <td rowspan="1" colspan="1">Conservation status of Mediterranean dune habitats assessed across multiple spatial scales using ecological and landscape indicators</td>
            <td rowspan="1" colspan="1">Medium (ecological surveys and spatial data)</td>
            <td rowspan="1" colspan="1">Mediterranean dunes</td>
            <td rowspan="1" colspan="1">EGDVI provides a fully standardised scoring system and explicit pressure and governance components</td>
            <td rowspan="1" colspan="1">(<xref ref-type="bibr" rid="B13882318">Carboni et al. 2009</xref>)</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">Vegetation-based conservation and pressure assessment</td>
            <td rowspan="1" colspan="1">Conservation status and anthropogenic impact assessed primarily through vegetation structure and composition</td>
            <td rowspan="1" colspan="1">Medium (vegetation and land-use data)</td>
            <td rowspan="1" colspan="1">Mediterranean dunes</td>
            <td rowspan="1" colspan="1">EGDVI strengthens geomorphological and legal dimensions absent from vegetation-centred approaches</td>
            <td rowspan="1" colspan="1">(<xref ref-type="bibr" rid="B13883003">Pinna et al. 2015</xref>)</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">Coastal / Mediterranean Dune Vulnerability Index (CDVI / MDVI)</td>
            <td rowspan="1" colspan="1">Physical, ecological and human variables (five components) scored on a 0–4 scale and averaged into vulnerability classes</td>
            <td rowspan="1" colspan="1">Medium (field surveys, GIS; equal weighting)</td>
            <td rowspan="1" colspan="1">Mediterranean dunes; tested in different regions</td>
            <td rowspan="1" colspan="1">EGDVI introduces weighted scoring and integrates scientific and legal value beyond vulnerability</td>
            <td rowspan="1" colspan="1">(<xref ref-type="bibr" rid="B13882388">Ciccarelli et al. 2017</xref>, <xref ref-type="bibr" rid="B13882287">Bertoni et al. 2019</xref>)</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">Vegetation-based Risk Index (VRI)</td>
            <td rowspan="1" colspan="1">Risk assessment combining hazard and vulnerability dimensions, including dune habitat and vegetation response to disturbance</td>
            <td rowspan="1" colspan="1">High (ecological, socio-economic and spatial datasets)</td>
            <td rowspan="1" colspan="1">Mediterranean dunes</td>
            <td rowspan="1" colspan="1">EGDVI focuses on eco-geomorphological value rather than risk and excludes socio-economic exposure metrics</td>
            <td rowspan="1" colspan="1">(<xref ref-type="bibr" rid="B13882223">Alessandrini et al. 2024</xref>)</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">StaDun / BeaPot / CoMan / SurLan composite indices + Dune Restoration Potential (DRP)</td>
            <td rowspan="1" colspan="1">Composite checklist indices assessing beach–dune system status, management pressure and restoration feasibility</td>
            <td rowspan="1" colspan="1">Medium–High (field surveys, orthophotos, management data)</td>
            <td rowspan="1" colspan="1">Mediterranean developed coasts</td>
            <td rowspan="1" colspan="1">EGDVI focuses exclusively on dune habitats and value assessment, avoiding mixing beach and dune processes</td>
            <td rowspan="1" colspan="1">(<xref ref-type="bibr" rid="B13882994">Pinna et al. 2022</xref>, <xref ref-type="bibr" rid="B13882456">Della Bella et al. 2024</xref>)</td>
          </tr>
          <tr>
            <td rowspan="1" colspan="1">Eco-Geomorphological Dune Value Index (EGDVI)</td>
            <td rowspan="1" colspan="1">Geomorphology, ecology, anthropogenic pressure, legal protection and scientific value (31 indicators; 0–4 scoring)</td>
            <td rowspan="1" colspan="1">Medium–High (field surveys, UAS/GIS, legal and scientific data)</td>
            <td rowspan="1" colspan="1">Black Sea coast; transferable to microtidal settings</td>
            <td rowspan="1" colspan="1">Introduced in this study</td>
            <td rowspan="1" colspan="1">This study (Prodanov et al. 2026)</td>
          </tr>
        </tbody>
      </table>
    </table-wrap>
    <supplementary-material id="S13883629" orientation="portrait" position="float" xlink:type="simple">
      <object-id content-type="arpha">96595C76-FF4D-509F-BC0B-999F366500C2</object-id>
      <object-id content-type="doi">10.3897/oneeco.11.e189660.suppl1</object-id>
      <label>Supplementary material 1</label>
      <caption>
        <p>Indicator definitions and scoring scheme for the EGDVI</p>
      </caption>
      <statement content-type="dataType">
        <label>Data type</label>
        <p>Description of indicator</p>
      </statement>
      <statement content-type="notes">
        <label>Brief description</label>
        <p>This supplementary file provides the full set of indicator definitions and the detailed scoring scheme used for the Eco-Geomorphological Dune Value Index (EGDVI). For each of the 31 indicators, grouped into eight thematic categories (geomorphology, aeolian dynamics, ecology, pressure, human footprint, legal protection, scientific value and nature conservation), qualitative criteria are defined for a standardised 0–4 scoring scale. The scheme ensures transparent, repeatable and objective evaluation of dune system condition and value and forms the methodological basis for indicator scoring and index calculation applied in the present study.</p>
      </statement>
      <p>File: oo_1635961.docx</p>
      <media xlink:href="oneeco-11-e189660-s001.docx" mimetype="Microsoft Word Document" mime-subtype="docx" position="float" orientation="portrait" xlink:type="simple">
        <uri content-type="original_file">https://binary.pensoft.net/file/1635961</uri>
      </media>
      <attrib specific-use="authors">Bogdan Prodanov, Radoslava Bekova, Magdalena Valcheva, Rosen Tzonev, Ahinora Baltakova</attrib>
    </supplementary-material>
    <supplementary-material id="S13886327" orientation="portrait" position="float" xlink:type="simple">
      <object-id content-type="arpha">76F536B3-890C-59A5-AD18-5F93FE4645CD</object-id>
      <object-id content-type="doi">10.3897/oneeco.11.e189660.suppl2</object-id>
      <label>Supplementary material 2</label>
      <caption>
        <p>Management framework for the EGDVI classes (A–E): definitions, conservation status, intent and recommended actions</p>
      </caption>
      <statement content-type="dataType">
        <label>Data type</label>
        <p>Management framework / conservation guidelines</p>
      </statement>
      <statement content-type="notes">
        <label>Brief description</label>
        <p>This supplementary file presents the management framework linked to the EGDVI classification system (Classes A–E). For each eco-geomorphological value class, it defines the conservation status, management intent and a set of recommended actions, ranging from strict protection of near-reference dune systems to emergency intervention and feasibility-led rehabilitation of degraded sites. The framework translates EGDVI class results into actionable guidance for conservation planning, restoration prioritisation and coastal management and supports the use of the index as a decision-support tool for policy and practice.</p>
      </statement>
      <p>File: oo_1635963.docx</p>
      <media xlink:href="oneeco-11-e189660-s002.docx" mimetype="Microsoft Word Document" mime-subtype="docx" position="float" orientation="portrait" xlink:type="simple">
        <uri content-type="original_file">https://binary.pensoft.net/file/1635963</uri>
      </media>
      <attrib specific-use="authors">Bogdan Prodanov, Radoslava Bekova, Magdalena Valcheva, Rosen Tzonev, Ahinora Baltakova</attrib>
    </supplementary-material>
    <supplementary-material id="S13886328" orientation="portrait" position="float" xlink:type="simple">
      <object-id content-type="arpha">050643E3-86B4-501D-B86F-34C2C7B6DC9A</object-id>
      <object-id content-type="doi">10.3897/oneeco.11.e189660.suppl3</object-id>
      <label>Supplementary material 3</label>
      <caption>
        <p>Floristic data from 16 dune systems along the Strandzha Black Sea Coast, Bulgaria</p>
      </caption>
      <statement content-type="dataType">
        <label>Data type</label>
        <p>Floristic data</p>
      </statement>
      <statement content-type="notes">
        <label>Brief description</label>
        <p>Species lists are composed of both original (collected May – September 2025) and published data (Gussev and Tzonev 2014; Valcheva et al. 2020, 2021) – 113 phytosociological plots (5 m × 5 m), of which 91 original, as well as additional floristic lists from the studied dune systems. Locations are numbered as follows: 1 – Primorsko (Stamopolu – Perla); 2 – Primorsko (International Youth Centre); 3 – Atliman; 4 – Kiten; 5 – Koral and Campsite Yug; 6 – Lozenets; 7 – Oazis; 8 – Arapya; 9 – Tsarevo (Popski plazh); 10 – Nestinarka; 11 – Ahtopol; 12 – Veleka River Mouth; 13 – Butamyata; 14 – Lipite; 15 – Listi; 16 – Silistar. Different species groups are coded as follows: P – psammophytes (pioneer dune species marked with red letter), W – weeds and ruderals, G – plants typical of grasslands and shrublands, F – plants typical of forests, A – alien plant species (species with invasive potential marked with red letter), B – bryophytes, L – lichenised fungi (assignment to species groups is according to their main habitat preference, using the approach of Valcheva et al. (2020). Vascular plants nomenclature was applied according to Euro+Med PlantBase (2006–2025). Bryophyte nomenclature was applied according to Hill et al. (2006) and nomenclature of lichenised fungi was applied according to Nimis and Martellos (2003).</p>
      </statement>
      <p>File: oo_1635964.docx</p>
      <media xlink:href="oneeco-11-e189660-s003.docx" mimetype="Microsoft Word Document" mime-subtype="docx" position="float" orientation="portrait" xlink:type="simple">
        <uri content-type="original_file">https://binary.pensoft.net/file/1635964</uri>
      </media>
      <attrib specific-use="authors">Bogdan Prodanov, Radoslava Bekova, Magdalena Valcheva, Rosen Tzonev, Ahinora Baltakova</attrib>
    </supplementary-material>
    <supplementary-material id="S13886329" orientation="portrait" position="float" xlink:type="simple">
      <object-id content-type="arpha">113780DE-3995-50FF-8A41-D48C6BA8134A</object-id>
      <object-id content-type="doi">10.3897/oneeco.11.e189660.suppl4</object-id>
      <label>Supplementary material 4</label>
      <caption>
        <p>Excel-based assessment tool for the Eco-Geomorphological Dune Value Index (EGDVI))</p>
      </caption>
      <statement content-type="dataType">
        <label>Data type</label>
        <p>Calculation tool/spreadsheet</p>
      </statement>
      <statement content-type="notes">
        <label>Brief description</label>
        <p>This supplementary file contains the Excel-based assessment tool used to calculate the Eco-Geomorphological Dune Value Index (EGDVI) for individual coastal dune systems. The tool implements indicator scoring, weighting, aggregation and class assignment for all 31 EGDVI indicators, grouped into eight thematic categories, using a standardised 0–4 scoring scale and expert-derived weights. It enables transparent, reproducible and automated calculation of total EGDVI scores and corresponding value classes (A–E) and was used for the pilot application to the Strandzha Black Sea Coast.</p>
      </statement>
      <p>File: oo_1635965.xlsx</p>
      <media xlink:href="oneeco-11-e189660-s004.xlsx" mimetype="Microsoft Excel Document" mime-subtype="xlsx" position="float" orientation="portrait" xlink:type="simple">
        <uri content-type="original_file">https://binary.pensoft.net/file/1635965</uri>
      </media>
      <attrib specific-use="authors">Bogdan Prodanov, Radoslava Bekova, Magdalena Valcheva, Rosen Tzonev, Ahinora Baltakova</attrib>
    </supplementary-material>
  </floats-group>
</article>
